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Encyclopedia of Evolution.pdf - Online Reading Center

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Gould, Stephen Jay<br />

animals, and the changes that occurred during the course<br />

<strong>of</strong> human evolution, it is necessary to thoroughly study the<br />

behavior <strong>of</strong> other animals, especially chimpanzees, under natural<br />

conditions. This required many years <strong>of</strong> close observation<br />

and the ability to recognize individual chimpanzees. Goodall’s<br />

close and prolonged observation yielded surprises that other<br />

researchers <strong>of</strong> animal behavior had not seen.<br />

Goodall was born April 3, 1934. In 1960 Goodall<br />

began observations in the Gombe Preserve in Tanzania.<br />

One persistent problem in the study <strong>of</strong> animal behavior<br />

(see behavior, evolution <strong>of</strong>) is that the behavior <strong>of</strong> the<br />

animals, particularly intelligent ones like chimpanzees, is<br />

altered by the presence <strong>of</strong> human observers. Months passed<br />

before the chimpanzees allowed Goodall to approach them<br />

closely enough for observation. Once they had accepted<br />

her, she was able to observe their normal lives. In her first<br />

year <strong>of</strong> observation Goodall made two discoveries. First,<br />

she found that chimpanzees hunt and eat meat. Second,<br />

she found that chimpanzees use tools. She observed chimps<br />

stripping leaves from twigs and using the twigs to fish termites<br />

out <strong>of</strong> a nest. This observation proved that humans<br />

were not the only animals that made and used tools (see<br />

technology). Goodall earned her Ph.D. in ethology from<br />

Cambridge University in 1965.<br />

Soon thereafter Goodall returned to Tanzania and<br />

established the Gombe Stream Research <strong>Center</strong>. She continued<br />

to make observations that showed chimpanzees to<br />

be capable <strong>of</strong> behavior previously associated only with<br />

humans. Particularly striking is the complexity <strong>of</strong> chimpanzee<br />

social organization and the advance planning that<br />

allows it. She observed a female chimpanzee adopt an unrelated<br />

baby chimpanzee. Chimpanzees were also capable<br />

<strong>of</strong> actions that remind observers <strong>of</strong> the bad side <strong>of</strong> human<br />

behavior. In 1974 a four-year war began between two<br />

bands <strong>of</strong> chimpanzees, resulting in the annihilation <strong>of</strong> one.<br />

Goodall also observed one chimpanzee that cannibalized<br />

several infants. Perhaps the most striking observation was<br />

in 1970 when Goodall observed chimpanzees apparently<br />

dancing for joy in front <strong>of</strong> a waterfall. All <strong>of</strong> these observations<br />

call into question the vast gulf that most people,<br />

including many scientists, assume to exist between humans<br />

and other animal species, and demonstrate that individual<br />

chimpanzees can differ as much from one another as individual<br />

humans.<br />

Goodall is interested in far more than just understanding<br />

animal behavior. She began and continues efforts to<br />

rescue orphaned chimpanzees and to improve the conditions<br />

<strong>of</strong> chimpanzees that are used in medical research.<br />

The president <strong>of</strong> a medical research company who at first<br />

despised Goodall’s interference ended up thanking her<br />

for her work. In 1977 Goodall founded the Jane Goodall<br />

Institute for Wildlife Research, Education, and Conservation,<br />

which encourages local people in Africa and all over<br />

the world to undertake conservation efforts in their own<br />

region. Her work both in the scientific study <strong>of</strong> behavior<br />

and in conservation have earned her many awards and<br />

worldwide recognition.<br />

Further <strong>Reading</strong><br />

Goodall, Jane, and Phillip Berman. Reason for Hope: A Spiritual<br />

Journey. New York: Warner Books, 1999.<br />

Jane Goodall Institute. “The Jane Goodall Institute.” Available online.<br />

URL: http://www.janegoodall.org. Accessed March 28, 2005.<br />

Miller, Peter. “Jane Goodall.” National Geographic, December 1995,<br />

102–129.<br />

Peterson, Dale. Jane Goodall: The Woman Who Redefined Man.<br />

New York: Houghton Mifflin, 2006.<br />

Gould, Stephen Jay (1941–2002) American <strong>Evolution</strong>ary<br />

biologist Stephen Jay Gould was one <strong>of</strong> the most creative<br />

thinkers in the study <strong>of</strong> evolution and one <strong>of</strong> the most famous<br />

and effective popularizers <strong>of</strong> evolution to the general public. His<br />

first scientific book, Ontogeny and Phylogeny, and his first popular<br />

book, Ever Since Darwin, were both published in 1977.<br />

Born September 10, 1941, Gould became interested in<br />

evolution when he was five years old and saw the Tyrannosaurus<br />

skeleton at the American Museum <strong>of</strong> Natural History<br />

in New York. He stayed with his youthful decision to become<br />

an evolutionary scientist. After graduating from Antioch College,<br />

Gould earned his Ph.D. from Columbia University. Until<br />

his death he was a pr<strong>of</strong>essor at Harvard University and a<br />

curator at Harvard’s Museum <strong>of</strong> Comparative Zoology.<br />

One <strong>of</strong> Gould’s insights into evolution was that evolution<br />

was not entirely a gradual process. Instead, periods <strong>of</strong><br />

rapid change, associated with the appearance <strong>of</strong> a new species<br />

(see speciation), were followed by long periods <strong>of</strong> stasis.<br />

Together with a colleague at the American Museum <strong>of</strong><br />

Natural History (see Eldredge, Niles), Gould presented the<br />

theory <strong>of</strong> punctuated equilibria in 1972. Gould, Eldredge,<br />

and Yale paleontologist Elisabeth Vrba maintained that evolution<br />

occurs not just on the level <strong>of</strong> populations (see natural<br />

selection) but on the species level as well. While defending<br />

the pro<strong>of</strong> <strong>of</strong> evolution and the mechanism <strong>of</strong> natural selection<br />

that Darwin had presented (see Darwin, Charles;<br />

origin <strong>of</strong> species [book]), Gould maintained that Darwin<br />

had unnecessarily constrained the operation <strong>of</strong> evolution by<br />

claiming that it always occurred gradually. Gould summarized<br />

a lifetime <strong>of</strong> insights into the processes <strong>of</strong> evolution in<br />

The Structure <strong>of</strong> <strong>Evolution</strong>ary Theory. Another major insight<br />

was that many features <strong>of</strong> organisms were not adaptations<br />

but were side effects <strong>of</strong> adaptations, resulting from structural<br />

constraints. Gould and paleontologist Elisabeth Vrba called<br />

these features exaptations (see adaptation).<br />

A related insight was that chance events have played a<br />

major role in the histories <strong>of</strong> individual humans, <strong>of</strong> societies,<br />

<strong>of</strong> species, and <strong>of</strong> the entire Earth. Gould’s doctoral research<br />

focused on land snails (genus Cerion) in Bermuda. He closely<br />

studied the individual variations among these snails, the same<br />

way that Darwin focused upon barnacles. Gould found that<br />

many <strong>of</strong> the variations could be better explained as the products<br />

<strong>of</strong> chance, or <strong>of</strong> structural constraint, than <strong>of</strong> adaptation.<br />

Gould also survived cancer for more than two decades. This<br />

experience contributed to his thinking about the importance<br />

<strong>of</strong> chance events. Gould’s 1989 book Wonderful Life made<br />

the fossils <strong>of</strong> the Burgess shale, from a 510-million-year-old

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