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Encyclopedia of Evolution.pdf - Online Reading Center

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• Proterozoic Eon (“earlier life”; also called Proterozoic<br />

era). During the Proterozoic, oxygen gradually accumulated<br />

in the atmosphere, and complex cells evolved (see<br />

eukaryotes, evolution <strong>of</strong>). In the middle <strong>of</strong> the Proterozoic,<br />

a few multicellular organisms such as seaweeds<br />

evolved. Toward the end <strong>of</strong> the Proterozoic, during the<br />

Ediacaran period, there were a few animals, as evidenced<br />

by embryos and burrows. Most <strong>of</strong> the inhabitants <strong>of</strong> the<br />

Ediacaran period were the enigmatic Ediacaran organisms.<br />

By the end <strong>of</strong> the Proterozoic, the atmosphere had<br />

its modern concentration <strong>of</strong> oxygen. At least two worldwide<br />

glaciations occurred during the Proterozoic (see<br />

Snowball Earth). The Ediacaran period (defined in<br />

2004) is the first new geological period to be defined in<br />

over a century. It lasted from about 600 to about 540 million<br />

years ago.<br />

All <strong>of</strong> Earth history after the Precambrian comprises the<br />

Phanerozoic Eon (“visible life”). This eon, which consists <strong>of</strong><br />

three eras, includes the evolution <strong>of</strong> all complex organisms<br />

that were related to modern plant and animal groups:<br />

• Paleozoic era (“ancient life”). Animal life proliferated in the<br />

oceans (see invertebrates, evolution <strong>of</strong>; fishes, evolution<br />

<strong>of</strong>). Land plants and animals evolved (see seedless<br />

plants, evolution <strong>of</strong>; gymnosperms, evolution <strong>of</strong>;<br />

amphibians, evolution <strong>of</strong>; reptiles, evolution <strong>of</strong>). The<br />

land had been barren at the beginning <strong>of</strong> the Paleozoic era<br />

but by the end was covered with extensive forests, which<br />

had modern groups <strong>of</strong> plants except the flowering plants.<br />

The end <strong>of</strong> the Paleozoic era was marked by the greatest <strong>of</strong><br />

the mass extinctions, in which over 95 percent <strong>of</strong> the species<br />

died (see Permian extinction). In most major groups<br />

<strong>of</strong> organisms, a few representatives survived this extinction.<br />

• Mesozoic era (“middle life”). Life proliferated in the<br />

oceans. On land, flowering plants evolved (see angiosperms,<br />

evolution <strong>of</strong>), and dinosaurs were the largest<br />

and most diverse <strong>of</strong> the vertebrates. Mammals and birds<br />

also evolved during the Mesozoic era (see birds, evolution<br />

<strong>of</strong>; mammals, evolution <strong>of</strong>). The Mesozoic era<br />

ended when a gigantic asteroid hit the Earth (see Cretaceous<br />

extinction).<br />

• Cenozoic era (“recent life”). A cooler, drier Earth was<br />

dominated by flowering plants, and the mammals were the<br />

largest vertebrates. Toward the end <strong>of</strong> the Cenozoic era,<br />

particularly cool, dry conditions caused the ice ages.<br />

Eras (particularly the last three) are divided into periods,<br />

most <strong>of</strong> which also reflect major events in Earth history. All<br />

<strong>of</strong> the periods are divided into epochs, although most scientists<br />

are familiar only with the epochs <strong>of</strong> the Tertiary and<br />

Quaternary periods. Because <strong>of</strong> the intense interest <strong>of</strong> humans<br />

in their own history, the Recent epoch has been defined to<br />

begin just 10,000 years ago, coinciding with the end <strong>of</strong> the<br />

most recent ice age and the beginning <strong>of</strong> villages, agriculture,<br />

and then civilization. The Earth is currently in the Phanerozoic<br />

Eon, the Cenozoic era, the Quaternary period, and the<br />

Recent epoch.<br />

The <strong>Encyclopedia</strong> <strong>of</strong> <strong>Evolution</strong> contains entries for each<br />

<strong>of</strong> the eras and periods <strong>of</strong> the Phanerozoic Eon. The epochs<br />

Goodall, Jane<br />

Geological Time Scale with Selected Periods<br />

and Epochs (to nearest million years)<br />

Duration<br />

(million<br />

Eons Eras Periods Epochs years ago)<br />

Hadean 4500–3800<br />

Archaean 3800–2500<br />

Proterozoic 2500–540<br />

Ediacaran 600–540<br />

Phanerozoic 540–<br />

Paleozoic 540–250<br />

Cambrian 540–510<br />

Ordovician 510–440<br />

Silurian 440–410<br />

Devonian 410–360<br />

Carboniferous 360–290<br />

Permian 290–250<br />

Mesozoic 250–65<br />

Triassic 250–210<br />

Jurassic 210–140<br />

Cretaceous 140–65<br />

Cenozoic 65–<br />

Tertiary 65–2<br />

Paleocene 65–55<br />

Eocene 55–35<br />

Oligocene 35–25<br />

Miocene 25–5<br />

Pliocene 5–2<br />

Quaternary 2–<br />

Pleistocene 2–0.01<br />

Holocene 0.01–<br />

Note: The complete list <strong>of</strong> periods is presented only for the Phanerozoic<br />

Eon.<br />

Note: The complete list <strong>of</strong> epochs is presented only for the Cenozoic era.<br />

Note: The Holocene epoch is also called the Recent epoch.<br />

Note: The Cenozoic era is now frequently divided into the Paleogene<br />

period (Paleocene, Eocene, Oligocene epochs) and the Neogene<br />

period (Miocene, Pliocene, Pleistocene, and Holocene epochs).<br />

<strong>of</strong> the Tertiary and Quaternary periods are included with<br />

their respective periods. In the accompanying table, as well<br />

as in each entry, the time periods have been rounded to the<br />

nearest five million years.<br />

Further <strong>Reading</strong><br />

Museum <strong>of</strong> Paleontology, University <strong>of</strong> California, Berkeley. “The<br />

paleontology portal.” Available online. URL: http://www.<br />

paleoportal.org/. Accessed May 3, 2005.<br />

Goodall, Jane (1934– ) British Primatologist Jane Goodall<br />

is a leading world authority on the species that is one <strong>of</strong><br />

humankind’s closest relatives, the chimpanzee (see primates).<br />

In order to understand how humans are different from other

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