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Encyclopedia of Evolution.pdf - Online Reading Center

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Skeleton <strong>of</strong> “Lucy,” the Australopithecus afarensis discovered by Donald<br />

Johanson and associates. It is one <strong>of</strong> the most complete australopithecine<br />

skeletons that has been found, and it demonstrated that upright posture<br />

preceded increase in brain size in human evolution. (Courtesy <strong>of</strong> Science<br />

VU/Visuals Unlimited)<br />

and the pelvis was suitable for walking. The feet, however,<br />

appear to have been only partly adapted to upright walking:<br />

the big toe was still at a noticeable angle. This suggests<br />

that Australopithecus lived mostly on the ground, walking<br />

upright, but frequently scrambled back into the trees when<br />

danger threatened. The relatively long arms and short legs<br />

<strong>of</strong> A. afarensis further suggest a mixture between arboreal<br />

australopithecines<br />

and ground life. It even suggests the possibility that upright<br />

posture began as an adaptation for shinnying up trees from<br />

brief visits to the ground and later allowed full adaptation<br />

to ground life.<br />

The primarily upright locomotion <strong>of</strong> A. afarensis and/<br />

or related australopithecines was confirmed by famous fossilized<br />

footprints found by anthropologist Mary Leakey (see<br />

Leakey, Mary) at Laetoli in Tanzania (see photo below).<br />

At about three million years <strong>of</strong> age, the footprints were<br />

undoubtedly produced by A. afarensis or a closely related<br />

species. Their date is fairly certain, as the footprints were<br />

imbedded in volcanic dust, on which radiometric dating<br />

can be used.<br />

Trail <strong>of</strong> hominin footprints in volcanic ash. This 00-foot ( 0-m) trail was<br />

discovered by Mary Leakey’s expedition at Laetoli, Tanzania, in . The<br />

footprints were probably made by Australopithecus afarensis individuals<br />

. million years ago. They demonstrate that hominins had already<br />

acquired the upright, free-striding gait <strong>of</strong> modern humans. The footprints<br />

have well-developed arches and the big toe does not diverge noticeably.<br />

They are <strong>of</strong> two adults with possibly a third set belonging to a child who<br />

walked in the footsteps <strong>of</strong> one <strong>of</strong> the adults. (Courtesy <strong>of</strong> John Reader/<br />

Photo Researchers, Inc.)

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