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Encyclopedia of Evolution.pdf - Online Reading Center

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iodiversity<br />

Biodiversity (as indicated by the number <strong>of</strong> families <strong>of</strong> fossilized marine and terrestrial organisms) has increased through geological time. After each <strong>of</strong><br />

the five mass extinctions in earth history (five arrows), biodiversity decreased (especially after the Permian extinction, largest arrow) but within a few<br />

million years began to recover and continue its increase. (Redrawn from Benton)<br />

abundant photosynthetic organism in the oceans, was not discovered<br />

until 1988. There is no end in sight for discovering<br />

new insect species. Partly this is because there are so many<br />

<strong>of</strong> them (350,000 species <strong>of</strong> beetles and counting!) but also<br />

because they are so hard to see.<br />

Only an expert can recognize many <strong>of</strong> the distinctions<br />

between closely related species. Flies are <strong>of</strong>ten classified using<br />

the arcane science <strong>of</strong> chaetotaxy, which distinguishes them<br />

on the basis <strong>of</strong> the arrangements <strong>of</strong> their bristles. Clearly, one<br />

limiting factor to the discovery <strong>of</strong> new species is the availability<br />

<strong>of</strong> experts who can recognize that they are new.<br />

About a million and a half species have been named.<br />

Some biologists estimate that there may be as many as 30<br />

million to 100 million species in the world. Ecologist Terry<br />

Erwin calculated this estimate. He began by using pesticides<br />

to kill all the insects in certain tropical trees and collecting the<br />

insects that fell. By estimating how many <strong>of</strong> these insects were<br />

unique to certain species <strong>of</strong> trees, and extrapolating to the<br />

number <strong>of</strong> tree species, he was able to estimate the number <strong>of</strong><br />

tropical insect species. From there he could estimate the number<br />

<strong>of</strong> insect species in the world, and then <strong>of</strong> all other species,<br />

using existing proportions <strong>of</strong> each taxonomic category.<br />

Although his estimate is probably high, it is certain that there<br />

are many more species than those already discovered. Robert<br />

May estimates just under seven million (see table at right).<br />

Estimate <strong>of</strong> Global Species Diversity<br />

(Excluding Prokaryotes)<br />

Taxon Estimated number <strong>of</strong> species<br />

Protist kingdom 400,000<br />

Plant kingdom 320,000<br />

Fungus kingdom 500,000<br />

Animal kingdom<br />

Nematodes 500,000<br />

Molluscs 120,000<br />

Arthropods<br />

Crustaceans 150,000<br />

Arachnids 500,000<br />

Insects 4,000,000<br />

Vertebrates<br />

Fishes 27,000<br />

Amphibians 4,000<br />

Reptiles 7,150<br />

Birds 9,800<br />

Mammals 4,800<br />

Others 150,000<br />

Total 6,790,000

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