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Encyclopedia of Evolution.pdf - Online Reading Center

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Marine life. All modern groups <strong>of</strong> marine organisms<br />

existed during the Carboniferous, except aquatic mammals.<br />

Many older lineages <strong>of</strong> fishes, such as the armored fishes <strong>of</strong><br />

the preceding Devonian period, became extinct, while the<br />

fish lineages that are dominant today proliferated (see fishes,<br />

evolution <strong>of</strong>).<br />

Life on land. Extensive swamps filled the shallow seas<br />

and lakes <strong>of</strong> the Carboniferous period in the tropical areas.<br />

Plant and animal life included:<br />

• Plants. Large trees had been rare in the Devonian period,<br />

but trees up to 100 feet (30 m) in height became abundant<br />

in the Carboniferous. Most <strong>of</strong> these were tree-sized versions<br />

<strong>of</strong> modern club mosses and horsetails (see seedless<br />

plants, evolution <strong>of</strong>). These trees had systems <strong>of</strong> water<br />

transport and sexual reproduction that would not work<br />

very well for tree-sized plants on the Earth today but functioned<br />

well in the very wet conditions <strong>of</strong> the Carboniferous.<br />

The first seed plants evolved, possibly in drier upland<br />

regions, during this time (see gymnosperms, evolution<br />

<strong>of</strong>). The seed plants became more common, and the seedless<br />

trees began their slide toward extinction, during a period<br />

<strong>of</strong> drier weather during the Carboniferous. Some seed<br />

plants, such as the seed ferns, were in evolutionary lineages<br />

that are now extinct. There were no flowering plants.<br />

• Animals. Insect groups such as dragonflies and beetles were<br />

abundant during this period. Many <strong>of</strong> them, particularly<br />

dragonflies, were larger than any insects now alive on the<br />

Earth. Insects breathe through openings along their abdomens,<br />

which is an efficient method <strong>of</strong> gas exchange only in<br />

small animals or under conditions <strong>of</strong> elevated oxygen concentration.<br />

The large size <strong>of</strong> some Carboniferous insects has<br />

suggested to some paleontologists that the amount <strong>of</strong> oxygen<br />

in the atmosphere greatly exceeded that <strong>of</strong> today. The<br />

insects were predators or ate plant spores; herbivores, that<br />

ate leaf material, were apparently very rare. The shortage <strong>of</strong><br />

herbivores may have contributed to the great accumulation<br />

<strong>of</strong> plant material into what became coal. Amphibians had<br />

evolved during the preceding Devonian period (see amphibians,<br />

evolution <strong>of</strong>) and proliferated during the Carboniferous<br />

into many forms, some large. The first vertebrates with<br />

hard-shelled eggs, which would today be classified as reptiles,<br />

evolved during the Carboniferous period (see reptiles, evolution<br />

<strong>of</strong>). Although some reptile lineages are not known<br />

until the Permian period, all <strong>of</strong> the major reptile lineages had<br />

probably separated before the end <strong>of</strong> the Carboniferous period.<br />

There were not yet any dinosaurs, mammals, or birds.<br />

Extinctions. The Carboniferous period is not recognized<br />

to contain one <strong>of</strong> the major extinction events. A worldwide<br />

period <strong>of</strong> warmer, drier climate near the end <strong>of</strong> the Carboniferous<br />

created conditions that favored the spread and diversification<br />

<strong>of</strong> seed plants and reptiles.<br />

Further <strong>Reading</strong><br />

White, Toby, Renato Santos, et al. “The Carboniferous.” Available<br />

online. URL: http://www.palaeos.com/Paleozoic/Carboniferous/<br />

Carboniferous.htm. Accessed March 23, 2005.<br />

catastrophism<br />

catastrophism Catastrophism was a set <strong>of</strong> geological theories<br />

that claimed that Earth history was dominated by a series<br />

<strong>of</strong> worldwide catastrophes. This approach dominated geology<br />

through the early 19th century, before uniformitarianism<br />

gained prominence (see Hutton, James; Lyell, Charles).<br />

Catastrophists, among whom Georges Cuvier in France<br />

and William Buckland in England figured prominently (see<br />

Cuvier, Georges), believed that the Earth’s geological history<br />

was divided into a series <strong>of</strong> ages, separated by spectacular<br />

worldwide catastrophes. They believed that the Flood <strong>of</strong> Noah<br />

was the most recent <strong>of</strong> these catastrophes. Some modern creationists<br />

have called themselves catastrophists. They attribute<br />

all <strong>of</strong> the geological deposits to the Flood <strong>of</strong> Noah, on an Earth<br />

recently created. However, this position was not held among<br />

catastrophist geologists <strong>of</strong> the 19th century. It is primarily a<br />

product <strong>of</strong> 20th-century fundamentalism (see creationism).<br />

In studying the geological deposits <strong>of</strong> the Paris Basin,<br />

Cuvier determined that the Earth had experienced a long<br />

history, with many ages previous to this one. He found skeletons<br />

<strong>of</strong> mammals <strong>of</strong> kinds that no longer exist, from terrestrial<br />

deposits, with oceanic deposits over them; to him this<br />

meant that a catastrophic flood had destroyed the world in<br />

which those mammals lived. Each previous age <strong>of</strong> the Earth<br />

was destroyed by a flood, then repopulated with plants and<br />

animals created anew by the Creator. Each time, the world<br />

became more and more suitable for human habitation, which<br />

Cuvier thought was the Creator’s ultimate goal. The last<br />

<strong>of</strong> the floods, the Flood <strong>of</strong> Noah, was so sudden as to have<br />

left mammoths, hair and all, frozen in ice. As a result <strong>of</strong> the<br />

research <strong>of</strong> Louis Agassiz (see Agassiz, Louis), geologists<br />

had to admit that this most recent supposed flood was actually<br />

an ice age, in which glaciers, not floodwaters, had piled<br />

up moraines <strong>of</strong> rubble (see ice ages).<br />

Prior to the 19th century division between catastrophists<br />

and uniformitarians, the 18th century had been dominated<br />

by the Plutonists and the Neptunists. The Plutonists claimed<br />

that the geological history <strong>of</strong> the Earth had been dominated<br />

by volcanic eruptions and earthquakes. The Neptunists, in<br />

contrast, emphasized the role <strong>of</strong> worldwide floods. The Plutonists<br />

demanded to know, where did the water go after<br />

worldwide floods? The Neptunists demanded to know, without<br />

worldwide floods, how did fossilized seashells end up on<br />

mountaintops? Nineteenth-century catastrophists accepted<br />

both volcanic and flood processes, while uniformitarians<br />

rejected the worldwide effects <strong>of</strong> both processes. None <strong>of</strong><br />

them suspected that the continents might actually move (see<br />

continental drift).<br />

The teleological and supernaturalistic approach <strong>of</strong> catastrophism<br />

had to give way to Lyell’s scientific approach in<br />

order for progress to continue in geological sciences. Cuvier<br />

was not totally wrong nor Lyell totally right. The catastrophists<br />

believed in an Earth in which biological progress<br />

occurred, while Lyell believed in uniformity <strong>of</strong> state; the<br />

former approach opens the way to evolutionary science.<br />

Catastrophists, furthermore, realized that there were abrupt<br />

transitions between one geological age and another, while<br />

Lyell believed in uniformity <strong>of</strong> rate; the former approach

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