Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
Encyclopedia of Evolution.pdf - Online Reading Center
Create successful ePaper yourself
Turn your PDF publications into a flip-book with our unique Google optimized e-Paper software.
Marine life. All modern groups <strong>of</strong> marine organisms<br />
existed during the Carboniferous, except aquatic mammals.<br />
Many older lineages <strong>of</strong> fishes, such as the armored fishes <strong>of</strong><br />
the preceding Devonian period, became extinct, while the<br />
fish lineages that are dominant today proliferated (see fishes,<br />
evolution <strong>of</strong>).<br />
Life on land. Extensive swamps filled the shallow seas<br />
and lakes <strong>of</strong> the Carboniferous period in the tropical areas.<br />
Plant and animal life included:<br />
• Plants. Large trees had been rare in the Devonian period,<br />
but trees up to 100 feet (30 m) in height became abundant<br />
in the Carboniferous. Most <strong>of</strong> these were tree-sized versions<br />
<strong>of</strong> modern club mosses and horsetails (see seedless<br />
plants, evolution <strong>of</strong>). These trees had systems <strong>of</strong> water<br />
transport and sexual reproduction that would not work<br />
very well for tree-sized plants on the Earth today but functioned<br />
well in the very wet conditions <strong>of</strong> the Carboniferous.<br />
The first seed plants evolved, possibly in drier upland<br />
regions, during this time (see gymnosperms, evolution<br />
<strong>of</strong>). The seed plants became more common, and the seedless<br />
trees began their slide toward extinction, during a period<br />
<strong>of</strong> drier weather during the Carboniferous. Some seed<br />
plants, such as the seed ferns, were in evolutionary lineages<br />
that are now extinct. There were no flowering plants.<br />
• Animals. Insect groups such as dragonflies and beetles were<br />
abundant during this period. Many <strong>of</strong> them, particularly<br />
dragonflies, were larger than any insects now alive on the<br />
Earth. Insects breathe through openings along their abdomens,<br />
which is an efficient method <strong>of</strong> gas exchange only in<br />
small animals or under conditions <strong>of</strong> elevated oxygen concentration.<br />
The large size <strong>of</strong> some Carboniferous insects has<br />
suggested to some paleontologists that the amount <strong>of</strong> oxygen<br />
in the atmosphere greatly exceeded that <strong>of</strong> today. The<br />
insects were predators or ate plant spores; herbivores, that<br />
ate leaf material, were apparently very rare. The shortage <strong>of</strong><br />
herbivores may have contributed to the great accumulation<br />
<strong>of</strong> plant material into what became coal. Amphibians had<br />
evolved during the preceding Devonian period (see amphibians,<br />
evolution <strong>of</strong>) and proliferated during the Carboniferous<br />
into many forms, some large. The first vertebrates with<br />
hard-shelled eggs, which would today be classified as reptiles,<br />
evolved during the Carboniferous period (see reptiles, evolution<br />
<strong>of</strong>). Although some reptile lineages are not known<br />
until the Permian period, all <strong>of</strong> the major reptile lineages had<br />
probably separated before the end <strong>of</strong> the Carboniferous period.<br />
There were not yet any dinosaurs, mammals, or birds.<br />
Extinctions. The Carboniferous period is not recognized<br />
to contain one <strong>of</strong> the major extinction events. A worldwide<br />
period <strong>of</strong> warmer, drier climate near the end <strong>of</strong> the Carboniferous<br />
created conditions that favored the spread and diversification<br />
<strong>of</strong> seed plants and reptiles.<br />
Further <strong>Reading</strong><br />
White, Toby, Renato Santos, et al. “The Carboniferous.” Available<br />
online. URL: http://www.palaeos.com/Paleozoic/Carboniferous/<br />
Carboniferous.htm. Accessed March 23, 2005.<br />
catastrophism<br />
catastrophism Catastrophism was a set <strong>of</strong> geological theories<br />
that claimed that Earth history was dominated by a series<br />
<strong>of</strong> worldwide catastrophes. This approach dominated geology<br />
through the early 19th century, before uniformitarianism<br />
gained prominence (see Hutton, James; Lyell, Charles).<br />
Catastrophists, among whom Georges Cuvier in France<br />
and William Buckland in England figured prominently (see<br />
Cuvier, Georges), believed that the Earth’s geological history<br />
was divided into a series <strong>of</strong> ages, separated by spectacular<br />
worldwide catastrophes. They believed that the Flood <strong>of</strong> Noah<br />
was the most recent <strong>of</strong> these catastrophes. Some modern creationists<br />
have called themselves catastrophists. They attribute<br />
all <strong>of</strong> the geological deposits to the Flood <strong>of</strong> Noah, on an Earth<br />
recently created. However, this position was not held among<br />
catastrophist geologists <strong>of</strong> the 19th century. It is primarily a<br />
product <strong>of</strong> 20th-century fundamentalism (see creationism).<br />
In studying the geological deposits <strong>of</strong> the Paris Basin,<br />
Cuvier determined that the Earth had experienced a long<br />
history, with many ages previous to this one. He found skeletons<br />
<strong>of</strong> mammals <strong>of</strong> kinds that no longer exist, from terrestrial<br />
deposits, with oceanic deposits over them; to him this<br />
meant that a catastrophic flood had destroyed the world in<br />
which those mammals lived. Each previous age <strong>of</strong> the Earth<br />
was destroyed by a flood, then repopulated with plants and<br />
animals created anew by the Creator. Each time, the world<br />
became more and more suitable for human habitation, which<br />
Cuvier thought was the Creator’s ultimate goal. The last<br />
<strong>of</strong> the floods, the Flood <strong>of</strong> Noah, was so sudden as to have<br />
left mammoths, hair and all, frozen in ice. As a result <strong>of</strong> the<br />
research <strong>of</strong> Louis Agassiz (see Agassiz, Louis), geologists<br />
had to admit that this most recent supposed flood was actually<br />
an ice age, in which glaciers, not floodwaters, had piled<br />
up moraines <strong>of</strong> rubble (see ice ages).<br />
Prior to the 19th century division between catastrophists<br />
and uniformitarians, the 18th century had been dominated<br />
by the Plutonists and the Neptunists. The Plutonists claimed<br />
that the geological history <strong>of</strong> the Earth had been dominated<br />
by volcanic eruptions and earthquakes. The Neptunists, in<br />
contrast, emphasized the role <strong>of</strong> worldwide floods. The Plutonists<br />
demanded to know, where did the water go after<br />
worldwide floods? The Neptunists demanded to know, without<br />
worldwide floods, how did fossilized seashells end up on<br />
mountaintops? Nineteenth-century catastrophists accepted<br />
both volcanic and flood processes, while uniformitarians<br />
rejected the worldwide effects <strong>of</strong> both processes. None <strong>of</strong><br />
them suspected that the continents might actually move (see<br />
continental drift).<br />
The teleological and supernaturalistic approach <strong>of</strong> catastrophism<br />
had to give way to Lyell’s scientific approach in<br />
order for progress to continue in geological sciences. Cuvier<br />
was not totally wrong nor Lyell totally right. The catastrophists<br />
believed in an Earth in which biological progress<br />
occurred, while Lyell believed in uniformity <strong>of</strong> state; the<br />
former approach opens the way to evolutionary science.<br />
Catastrophists, furthermore, realized that there were abrupt<br />
transitions between one geological age and another, while<br />
Lyell believed in uniformity <strong>of</strong> rate; the former approach