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THE REPRODUCTIVE ELEMENTS OF PLANTS AND ANIMALS 97<br />
physical forces) in certain cases overcome gravitation equally with the vital forces, but it does not follow from this<br />
that gravitation as a power is extinguished. When gravitation (and this remark appUes to all the physical forces)<br />
is useful in building up the organism, it is employed by nature in conjunction with the vital forces ; and when<br />
capillarity, osmosis, &c., are more convenient, they are preferred. The vital and physical forces must be regarded<br />
in their totality, nor must the object in view be lost sight of. Natural laws and forces (vital, chemical, and<br />
physical) act and react upon each other ;<br />
and though at times some are inoperative, they are not annihilated, neither<br />
are they opposed when the final effect to be produced is taken into account. Organised structures are built up in<br />
accordance with natural laws. Thus, their parts and particles consist of substances which naturally have a<br />
chemical afiinity for each other ; those substances absorb and respire—that is, take in and give off fluids ; they<br />
exhibit, for the most part, capillary spaces and interspaces in their interior ; they have weight, porosity, elasticity,<br />
and various other purely physical properties. In this way chemical affinity, osmosis, capillarity, gravitation, &c.,<br />
have free play, and each in turn assists not only in forming and maintaining the body, but also in disintegrating<br />
it. The vital forces, if I may be allowed the expression, use and abuse the physical ones. They do not, they<br />
cannot, annihilate them. As well may we say the vital forces annihilate matter.<br />
THE REPRODUCTIVE ELEMENTS OF PLANTS AND ANIMALS ESSENTIALLY AND<br />
FUNDAMENTALLY DIFFER FROM THE BEGINNING<br />
Whatever may be said regarding the ultimate composition of protoplasm, the grand fact remains, that the<br />
germs and seeds of plants and the eggs of animals, apparently homogeneous and identical, invariably produce not<br />
only their own kind, but, during the process of development, substances which are altogether dissimilar. Even man,<br />
with his wealth of imagination and thought, is the product of an egg, which to the eye of the microscopist and<br />
chemist is, as to its several parts, homogeneous and identical. Yet this egg, with its several parts placed in exactly<br />
the same conditions, produces the tender pulp of the brain, the fiaccid muscles, the tough tendons, the elastic carti-<br />
lages, the hard bones, the stiU harder teeth, the enamel, a substance nearly as brittle as glass, and in due time the<br />
several secretions, the major portion of which is composed of water. To these are to be added the external skin,<br />
with its hair, nails, &c. ; the internal skin or mucous membrane, with its numerous glands ; the secreting and<br />
excreting organs ; the nerves ; the lymphatics ; and the blood-vessels, with their contained blood, so rich, so<br />
complex, so marvellous in composition. These various products, so diverse in form, composition, and function,<br />
are, we know, the product of an egg. Shall we be told, or, if told, shall we believe, that this egg is homogeneous<br />
and identical in all its parts and particles ? Identical these parts may be to the eye of sense, but to the eye of<br />
reason they are not so. Matter similarly constituted, similarly disposed, and similarly conditioned—that is, placed<br />
under precisely similar circumstances—behaves in precisely the same manner ; in other words, all the parts are<br />
infiuenced afike. If fife had to operate on absolutely identical matter, the living mass would be simply homogeneous<br />
that is, undifferentiated ; and if undifferentiated, it would have no power to build up either a plant or an animal.<br />
It is impossible to produce muscle, bone, brain, &c., from matter identical in ultimate composition. In the germ,<br />
the seed, the egg, even from the first, there is differentiation, actual or potential.-' The first embryonic changes<br />
are the result of this differentiation. It is this which enables the different parts of the embryo to attract and<br />
assinailate different substances and to build up different tissues. One embryonic change involves another, and<br />
gradually the individual is developed, each according to its kind. The weaver may not produce a web dissimilar in<br />
texture and pattern in the absence of dissimilar elements. He must be supplied with the warp, the woof, and the<br />
varied pigment. A queen-bee may be formed from an ordinary bee grub by changing the quantity and quality<br />
of its food, but no amount of change of diet can produce from the same grub a dragon-fly or a locust.<br />
In making these statements regarding the non-identity of protoplasm, bioplasm, and the several parts of<br />
^ Mr. Herbert Spencer, in his " Principles of Biology," expressed the belief that in reproduction the cell must be regarded as the morpho-<br />
logical unit, but that in addition there must be a chemical unit consisting of the simplest combination of molecules capable of displaying life.<br />
Others, who regarded the nucleus as the most important part of the cell, discovered that by the employment of suitable dyes one portion of the<br />
nucleus could he distinguished from all the other parts. This they designated the chromatin. The chromatin, it was found, could he broken up into<br />
a number of rod-like bodies, which were designated chromosmies. The chromosomes, when stained, were found in turn to be compound and<br />
divisible into microsomes. This much could be demonstrated by the aid of the microscope with stained nuclei : when its powers failed, Professor<br />
Weismann, by a process of reasoning, inferred that each of the microsomes consisted of a number of still smaller bodies, which he called determinants.<br />
The determinants Weismann regarded as the structures forming the germ-plasm which decided the nature and character of the adult organism.<br />
Weismann went further, and assumed that not even the determinants constituted the ultimate living elements. The detei-minants, he believed,<br />
were further resolvable into biophores. Weismann, who has done much to expiscate this most involved and diihcult subject, was by no means the<br />
first to deal with it in a more or less exhaustive manner. It has been a veritable bone of contention for over thirty years—each author employing<br />
his own nomenclature.<br />
From the foregoing it will be seen that the reproductive cell is by no means a simple structure, and that it is seen to be more complex in<br />
proportion as our opportunities gf investigating it inoreas?,<br />
VOL. I.<br />
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