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THE TRAVELLING ORGANS OF ANIMALS 219<br />
land. The same is true of the Umbs and small feet of the ostrich. In the ostrich the digits of each foot are<br />
reduced to t'.vo ; one digit being mainly employed in rurming.<br />
Five-toed typical mammals, with heavy, powerful frames, are not so well adapted for running on hard<br />
surfaces as the horses, but they still exist—as witness the elephant. There was no necessity to manufacture the<br />
horse from a five-toed mammal. The modified hmbs and sohd hoofs of the horse are rather to be regarded as a<br />
proof of type and of adaptation and design, and, as such, are to be placed in the same category as the swimming<br />
tail of the fish and reptile ; the swimming feet of the frog, beaver, ornithorhynchus, bird, seal, and walrus ; and<br />
the flying wings of the insect, bird, and bat. These are all examples of structures adapted to the media the animals<br />
are to traverse (Plates 1., li., and lii.). They are special creations rather than mere modifications. Environment<br />
and externahties cannot produce them : they are constructed in young animals during development and before<br />
they are required. They are means to ends, but do not prove consanguinity and descent.<br />
The rehabilitation of so-called missing links, and the modifications of existing structures, especially limb<br />
structures, do not afford a safe basis for constructing a chart of consanguinity and descent. The presence of teeth<br />
in the heads of a few extinct birds, the pattern of the teeth of the horse, and the decrease or increase in size, and<br />
the appearance or disappearance of certain limb bones in the same or other individuals, do not prove evolution,<br />
descent, or blood relationship. All these phenomena can be explained outside evolution and descent by reference<br />
to the requirements of hfe and the exigencies of locomotion. That a few extinct birds had teeth does not prove<br />
the descent of the bird from the reptile ;<br />
size of others give a vahd clue to the descent of the horse.<br />
neither do the suppression of certain limb bones and the greatly increased<br />
As a matter of fact, the changes and modifications introduced into the animal series, because of modifications<br />
in their organs of locomotion, are much greater than the data usually relied on for proving the doctrines of evolu-<br />
tion and descent. I have, therefore, to express my conviction that evolutionists are not on safe ground when they<br />
confine proofs of their views to one or two Unes of demonstration ; these being by no means the most obvious or<br />
convincing.<br />
Professor Huxley regards America as the original home of the horse, rather than Europe, and mentions inci-<br />
dentally that the animal died out on the American continent and had to be re-introduced. If this view be correct,<br />
it is difficult to understand how such a contingency could have arisen, considering the vast extent and the resources<br />
of the continent occupied. The dying out view is essentially opposed to the theory of environment.<br />
The evolution of the foot of the horse from a five-toed mammal is on a level with the evolution of the wing<br />
of the pterodactyl, bird, and bat, from some five-fingered unknown flying ancestor. The wing of the bat (Fig. 38)<br />
is composed of an arm, fore arm, wrist and five digits, with an outstretched membrane. The digits are nearly<br />
of the same length, and take an equal share in supporting the membrane, and in the function of flight. In the<br />
hand and phalanges of the pterodactyl (Fig. 37), as in the foot and phalanges of the horse, several of the digits<br />
are dwarfed, one of them (the fourth) being enormously developed, and forming the chief organ of locomotion. Cer-<br />
tain of the digits are dwarfed and rudimentary, but no one would, on this account, say that the ancient ptero-<br />
dactyl was evolved from some extinct animal resembling the modern bat, which is a more highly differentiated<br />
animal. Similar remarks are to be made of the wing of the bird. The wing of the bird is composed of a humerus<br />
(arm bone), a radius and ulna (fore arm bones), modified carpal or wrist bones, and several digits or finger bones<br />
run together and fused (Fig. 4:2). The arm and fore arm bones are independent and separate as in the primates<br />
and in ourselves. The wrist and finger bones, on the contrary, are modified almost beyond recognition ; certain<br />
of them being suppressed, while others are fused and sohdified to form one strong compound bone, which bears<br />
the great primary feathers, the most essential structures in flight. The very remarkable modifications occurring<br />
in the wing of the bird are quite as extraordinary as those occurring in the limb of the horse, but no one would<br />
be so rash as to suppose that the bird, because of certain modifications in its wing, was descended from any form<br />
of mammal, with distinct wrist bones and five distinct digits. Neither would the bat (a mammal, with five digits<br />
in its wing) furnish the ancestor of the pterodactyl (Fig. 37). The dropping of certain bones, and the enlarging and<br />
fusing of others, in the hmbs and travelhng organs of animals afford no proof of evolution in the strict or proper<br />
sense. Evolution involves differentiation and increase in the number of parts. It is opposed to devolution, or the<br />
decrease and suppression of parts. The increase and decrease of parts here referred to, which occur even in<br />
the embryo, while inimical to the doctrine of evolution, supply strong arguments in favour of type, a First Cause,<br />
design, and adaptation to given ends.<br />
The modifications required for locomotion can be traced in all animals, however diverse. They are not confined<br />
to alHed animals, and consequently, and strictly speaking, have nothing to do with evolution ;<br />
connecting hnks, and points of resemblance and continuity, in the evolving forms.<br />
evolution requiring<br />
The connecting hnks in locomotion do not support the theory of evolution'. Thus the galeopithecus or flying