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Picture - Cosmic Polymath

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DEVELOPMENT OF EMBRYONIC MEMBRANES OR LAYERS 389<br />

From the mesohlast proceed—<br />

(h) The generative and urinary organs (the epithelium of the bladder and urethra excepted).<br />

(i) The voluntary and involuntary muscles (the muscular fibres of the sweat glands excepted).<br />

(/) The vascular and lymphatic systems, serous membranes, and spleen.<br />

(k) The skeleton and connective tissues.<br />

From the hypoblast proceed—<br />

(I) The epithelium of the alimentary canal and the glands which open into it.<br />

(to) The epithelium of the urinary bladder.<br />

(w) The epitheUum of the Eustachian tube, the tympanum, the bronchial tubes, and the air sacs of the lungs.<br />

(o) The epithelium Uning the vesicles of the thyroid body, and the epitheUal nests of the thymus.<br />

Home investigators, following His, have still further divided the blastoderm. They describe the true mesoderm<br />

as consisting of two epitheUal lamellae. According to them the complete blastoderm would consist of four layers ;<br />

namely, epiblast, outer or somatic mesoblast, inner or splanchnic mesoblast, and hypoblast.<br />

The separation of the blastoderm into layers is followed by other changes in rapid succession. Very soon blood-<br />

vessels and a rudimentary heart, composed of nucleated cells (minus muscular fibres and nerves) make their appear-<br />

ance (Plate xc. Figs. 1, 3, 4, and 5). The heart, as already stated, is formed prior to its containing blood. The organ<br />

is made before it is called upon to discharge its peculiar function. The same holds true of other organs—the lungs,<br />

for example. The formation of organs before they are called upon to discharge specific functions affords the strongest<br />

possible evidence of design. This evidence is strengthened when it is borne in mind that the organs and systems are,<br />

in many cases, simultaneously produced. It is not a mere case of successive developments, but of successive and<br />

synchronous, continuous developments ;<br />

the Master-Architect and Builder carrying out His work in part piecemeal<br />

and in part by simultaneous, complicated, combined operations.<br />

The heart is not only made before it contains blood, but it also assumes peristaltic rhythmic movements for<br />

propelling the blood when it reaches its cavities. The blood, therefore, is not the cause of the movements of the heart,<br />

as is generally beUeved. As a matter of fact, the heart, before it contains either muscular fibres, nerves, or blood,<br />

opens and closes with time-regulated beat, and quite independently. The same holds true of the so-called contractile<br />

vesicles in plants and animals. This is due to the circumstance that the heart is a containing as well as a propelling<br />

and transmitting organ. All the hollow viscera (stomach, intestine, bladder, uterus, &c.), even in the adult, are<br />

endowed with similar powers. They are all containing as well as propelling organs.<br />

The changes which occur in the heart and vascular system during the process of development and in the transi-<br />

tion from the foetal to the adult state are of the most complicated and startling description, and will be adverted to<br />

further on when speaking of the placental, foetal, and adult circulations in man.<br />

The developing chick lives upon the albuminous and other materials in its vicinity, plus a little moisture and<br />

air. Organ after organ appears in anticipation of the function to be discharged by it. The brain and nerves,<br />

alimentary canal, glands, lungs, lymphatics, muscles, bones, feathers, &c., are all developed in a comparatively<br />

short period, and in advance of the peculiar work to be performed by each. The incubating process over, the<br />

completed bird emerges from what is practically an empty shell into the outer world, and immediately sets about<br />

procuring extraneous food.<br />

If the development of the chick within the egg is marvellous, and a triumph of design infinitely greater than<br />

that achieved by the watchmaker when he laboriously constructs a watch, the advent of the fully-fledged chick on<br />

terra firma and its instant and eager pursuit of food are, in a sense, still more wonderful. What an extraordinary<br />

combination of foresight and power does the development and birth of a chick and its immediate search for food as<br />

an independent being imply. All the functions of its multiple existence are at once brought into play : its lungs<br />

breathe air, its alimentary canal digests food, its muscular and nervous systems enable it to run here, there, and<br />

everywhere, its blood circulates, its glands secrete and excrete, and its brain and sense organs connect it in the most<br />

effective way with the outer world, in which it is henceforth to hve and move. Shall we be told, or if told, shall we<br />

believe, that the fully-developed living chick is the result of accident—a mere matter of spontaneous generation—<br />

product of irritability and artificial stimulation, or forsooth of environment ?

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