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Picture - Cosmic Polymath

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AMCEBA PROTEUS 333<br />

The amcsba draws in its pseudopodia and assumes a spherical shape at a temperature of 35' C. If the<br />

temperature be raised to 40° C. it dies. Under unfavourable circumstances, such as drought, the ectoplasm secretes<br />

a thin hyaline case, and the amcEba becomes encysted and dormant, in which condition it is carried by the wind or<br />

other means to various localities.<br />

The amoeba is invested with practically universal powers, and furnishes one of the most remarkable examples<br />

known of the potentialities residing in rudimentary living matter. Thus any part of its substance may become a<br />

stomach and digest its food, and any part of its body may throw off the waste products ; it has, however, neither<br />

mouth nor anus. It feeds upon diatoms, desmids, spores of algas, and other vegetable matter, and occasionally on<br />

animal matter, such as portions of rotifers and of Protozoa, notably Arcella.<br />

When the amoeba falls in with a microscopic alga or food particle, the protoplasm flows round it ; the ectoplasm<br />

opening to receive it and closing behind it. It enters the endoplasm with a drop of water which constitutes the<br />

food-vacuole, and is temporarily included in the body of the amoeba. The alga is slowly digested, its insoluble<br />

proteids being converted into soluble and diffusible proteids, which pass into the substance of the endoplasm. It<br />

Fig. 68.<br />

Fig. 68. Ama'ha pwteux as seen under varying conditions of size and shape (A to J), magnified from 400 to 800 diameters<br />

(after Masterman, and Marshall and Hurst).<br />

A. Amoeba in the encysted state, composed mainly of granular and clear sarcode ; c, contractile vesicle ; d, nucleus. It contains<br />

an oval-shaped diatom.<br />

B to J. Different shapes assumed by larger and smaller amoeba. In these figures the contractile vesicle is seen at c, the<br />

nucleus at d, and the finger-like projections at e. The projections, sometimes designated pseudopodia, are for the most part<br />

conicalin shape, as seen at C, D, E, F, G, H, and 1. They are caused by the animal voluntarily forging a part of its body outwards<br />

from the centre centrifugally. They are retracted in an opposite direction or centripetally. They are not due to contractions ; if<br />

they were, the projections would not be conical-shaped, but constricted where united to the body. In reality, they are produced<br />

by an aggressive, streaming, rhythmic movement similar to what occurs in the plasmodium of the Mycetozoa and gromia already<br />

described.<br />

At E, the body (a) of an amoeba is stretched over a large diatom (//).<br />

At J, the varying forms assumed by a small amoeba, at intervals of half-a-minute, are given.<br />

may be well to state that the proteids are themselves constituents of protoplasm. The amoeba, it is believed, cannot<br />

digest carbo-hydrates or fats, and does not build up its protoplasm from lower chemical constituents. The<br />

indigestible cellulose walls of alga and other intractable materials are extruded from the body. The amoeba<br />

inhales oxygen and exhales carbonic acid, in which respect it resembles the higher animal forms. It has, however,<br />

no branchiae, gills, or lungs ; its whole substance performing this important function.<br />

The reproduction of the amoeba is effected in the simplest manner possible. The nucleus, and with it the<br />

sarcode of the body, divides into two. In this respect the process resembles the first step in the segmentation of<br />

the impregnated ovum of the higher animals. There are, however (so far as known), no male and female elements,<br />

and the animal is not hermaphrodite. The original individual becomes two separate entities, each of which, after<br />

a short time, becomes as perfect as the parent, and so hfe is transmitted and continued in the most direct and effective<br />

manner. The reproduction of the amoeba is fundamental and thorough-going. It does not require two separate<br />

organs or individuals for its consummation. It is thought by some that the amoeba has not only the power of<br />

dividing into two by binary fission, but that two amoebae can come together, coalesce, and conjugate to form one<br />

individual. The process of conjugation, while not quite made out in the amoeba, certainly occurs in other low animal<br />

forms, such as the Gregarines and Infusoria. The powers inhering in what is practically undifferentiated living<br />

matter are at once striking and impressive. They bring us face to face with the eternal verities of matter and

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