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198 DESIGN IN NATURE<br />
processes of inorganic nature with the mental hfe of the highest animals. For all these phenomena—indeed, for<br />
all phenomena both in nature and in the mind—Lamarck takes exclusively mechanical, physical, and chemical<br />
activities to be the true efficient causes. Darwin refers them to natural selection. . . . All the phenomena of<br />
the psychic life are, without exception, bound up with certain material changes in the living substance of the body,<br />
the frotoflasm. We have given to that part of the protoplasm which seems to be the indispensable substratum<br />
of psychic life the name of fsychoflasm (the soul-substance in the monistic sense) ; in other words, we do not<br />
attribute any peculiar ' essence ' to it, but we consider the psyche to be merely a collective idea of all the psychic<br />
functions of protoplasm. In this sense the ' soul ' is merely a physiological abstraction like ' assimilation ' or<br />
' generation.' . . In the unicellular protista, the psychoplasm is identified either with the whole of the living<br />
protoplasm of the simple cell or with a portion of it. In all cases, in the lowest as well as the highest stages of<br />
the psychological hierarchy, a certain chemical composition and a certain physical activity of the psychoplasm<br />
are indispensable before the ' soul ' can function or act. That is equally true of the elementary psychic function<br />
of the plasmatic sensation and movement of the protozoa, and of the complex functions of the sense-organs and<br />
the brain in the higher animals and man. The activity of the psychoplasm, which we call the ' soul,' is always<br />
connected with metabohsm. ... All living organisms without exception have the faculty of spontaneous movement,<br />
in contradistinction to the rigidity and inertia of imorganised substances (for example, crystals) ; in other words,<br />
certain changes of place of the particles occur in the hving psychoplasm from internal causes, which have their<br />
source in its own chemical composition. ... At the lowest stage of organisation, in the lowest protists, the<br />
stimuli of the outer world (heat, hght, electricity, &c.), cause in the indifferent protoplasm only those indispensable<br />
movements of growth and nutrition which are common to all organisms and are absolutely necessary for their<br />
preservation. That is also the case in most of the plants. .<br />
. . The<br />
automatic and the reflex movements which<br />
we observe everywhere, even in the unicellular protists, seem to be the outcome of inchnations which are insepar-<br />
ably connected with the very idea of Hfe. Even in the plants and lowest animals these inchnations, or tropisms,<br />
seem to be the joint outcome of the inchnations of all the combined individual cells. When two cells meet as<br />
a result of copulation, or when they are brought into contact through artificial fertilisation] (in the fishes, for<br />
instance), they attract each other and become firmly attached. The main cause of this cellular attraction is a<br />
chemical sensitive action of the protoplasm, allied to smell or taste, which we call ' erotic chemicotropism ;<br />
' it may<br />
also be correctly (both in the chemical and the romantic sense) termed cellular afiinity or ' sexual cell-love.' On<br />
a critical study of these different embryonic formations, the evolution of which from each other we can directly<br />
observe under the microscope, we arrive, by means of the great law of biogeny, at a series of most important con-<br />
clusions as to the chief stages in the development of our psychic Hfe. We may distinguish (1) UniceUular protozoa<br />
with a simple cell-soul : the infusoria. (2) Multicellular protozoa with a communal soul : the catallacta. (3) The<br />
earliest metazoa with an epitheUal soul : the platodes. . . . The<br />
earliest ancestors of man and all other animals<br />
were unicellular protozoa. This fundamental hypothesis of rational phylogeny is based, in virtue of the phylo-<br />
genetic law, on the familiar embryological fact that every man, Hke every other metazoon (that is, every multi-<br />
cellular organism with tissues), begins his personal existence as a simple cell, the stem-cell (cytula), or the<br />
impregnated egg-cell. As this cell has a ' soul ' from the commencement, so had also the corresponding unicellular,<br />
ancestral forms, which were represented in the oldest series of man's ancestors by a number of different protozoa.<br />
. . . Every<br />
living cell has psychic properties, and the psychic Hfe of the multicellular animals and plants is merely<br />
the sum-total of the psychic functions of the cells which build up their structure. In the lower groups (in algse<br />
and sponges for instance) all the cells of the body have an equal share in it (or with very slight differences) ; in<br />
the higher groups, in harmony with the law of the ' division of labour,' only a select<br />
— ' the soul-cells.' . . . We<br />
portion of them are involved<br />
find the highest development of the animal cell-soul in the class of ciliata, or ciHated<br />
infusoria. The unicellular protozoa give proof of the possession of a highly-developed ' cell-soul,' which is of great<br />
interest for a correct decision as to the psyche of our earHest unicellular ancestors. . . . The<br />
tissue-soul (histo-<br />
psyche) : in all multicellular, tissue-forming plants (metaphyta) and in the lowest, nerveless classes of tissue-forming<br />
animals (metazoa) we have to distinguish two different forms of psychic activity—namely (1) the psyche of the<br />
individual ceHs which compose the tissue, and (2) the psyche of the tissue itself, or of the '<br />
cell-state ' which is<br />
made up of the tissues. This ' tissue-soul ' is the higher psychological function which gives physiological indi-<br />
viduaHty to the compound multicellular organism as a true ' cell-commonwealth.' It controls all the separate<br />
' cell-souls ' of the social cells—the mutually dependent ' citizens ' which constitute the community. .<br />
. . The<br />
plant-soul [phytopsyche) is, in our view, the summary of the entire psychic activity of the tissue-forming multi-<br />
cellular plant (the metaphyton, as distinct from the unicellular protophyton). .<br />
. The<br />
soul of the nerveless metazoa.<br />
Of very special interest for comparative psychology in general, and for the phylogeny of the animal soul in<br />
particular, is the psychic activity of those lower metazoa which have tissues, and sometimes differentiated organs.