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PREDICTIONS – 10 Years Later - Santa Fe Institute

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APPENDIX A<br />

Mathematical Formulation of S-Curves and the<br />

Procedure for Fitting Them on Data<br />

The behavior of populations growing under Darwinian competition is<br />

described in general by the celebrated Volterra-Lotka equations. These<br />

equations—one for every competing species—specify the rate of<br />

growth which is made up of several terms.<br />

dN i<br />

n<br />

—— = a i N i – ∑b ij N i N j<br />

dt j=1<br />

The positive term is proportional to the actual size. The parameter a i<br />

reflects the capacity of the species to reproduce, or, in other words, the<br />

rate of growth of population i in the absence of predators. The negative<br />

terms represent growth reduction due to loss of life caused by the predator.<br />

The parameter b ij can be interpreted as the rate at which predator j<br />

will grow assuming there is an infinite amount of prey i. The properties<br />

of the solutions to these equations have been described by Elliot Montroll<br />

and N. S. Goel 1 and more recently by M. Peschel and W. Mendel. 2<br />

We can distinguish the following special cases.<br />

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