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Evolution__3rd_Edition

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Glossary<br />

Words in italics cross-refer to a separate entry for that word<br />

elsewhere in the glossary.<br />

adaptation A feature of an organism enabling it to survive<br />

and reproduce in its natural environment better than if it lacked<br />

the feature.<br />

adaptive topography A graph of the average fitness of a<br />

population in relation to the frequencies of genotypes in it. Peaks<br />

on the landscape correspond to genotypic frequencies at which<br />

the average fitness is high, valleys correspond to genotypic frequencies<br />

at which the average fitness is low. It is also called an<br />

adaptive landscape, or a fitness surface.<br />

allele A variant of a single gene, inherited at a particular<br />

genetic locus; it is a particular sequence of nucleotides, coding for<br />

messenger RNA.<br />

allometry The relation between the size of an organism and<br />

the size of any of its parts: for example, there is an allometric<br />

relation between brain size and body size, such that (in this case)<br />

animals with bigger bodies have bigger brains. Allometric relations<br />

can be studied during the growth of a single organism,<br />

between different organisms within a species, or between organisms<br />

in different species.<br />

allopatric speciation Speciation via geographically separated<br />

populations.<br />

allopatry Living in separate places. Compare sympatry.<br />

amino acid A unit molecular building block of proteins. A protein<br />

is a chain of amino acids in a certain sequence. There are<br />

20 main amino acids in the proteins of living things, and the<br />

properties of a protein are determined by its particular amino<br />

acid sequence.<br />

amniotes The group comprising reptiles, birds, and mammals.<br />

They all develop through an embryo that is enclosed within<br />

a membrane called an amnion. The amnion surrounds the<br />

embryo with a watery substance, and is probably an adaptation<br />

for breeding on land.<br />

analogy A term mainly not used in this edition of the text, but<br />

close in meaning to homoplasy. That is, a character shared by<br />

a set of species but not present in their common ancestor a a<br />

convergently evolved character. Some biologists distinguish<br />

between homoplasies and analogies. In Chapter 3, the term is<br />

used to contrast with pre-evolutionary homology. An analogy is<br />

then a structure like a bird wing and an insect wing. It is similar<br />

for functional reasons and not deeply similar in structure.<br />

Compare homology.<br />

anatomy (i) The structure itself of an organism, or one of its<br />

parts. (ii) The science that studies those structures.<br />

ancestral homology Homology that evolved before the common<br />

ancestor of a set of species, and is present in other species<br />

outside that set of species. Compare derived homology.<br />

area cladogram A branching diagram (or phylogeny) of a set<br />

of species (or other taxa) showing the geographic areas they<br />

occupy. According to the theory of vicariance biogeography, the<br />

branching diagram represents the history of range splits (probably<br />

driven by geological processes such as continental drift) in<br />

the ancestry of the species.<br />

artificial selection Selective breeding, carried out by humans,<br />

to alter a population. The forms of most domesticated and<br />

agricultural species have been produced by artificial selection;<br />

it is also an important experimental technique for studying<br />

evolution.<br />

asexual reproduction The production of offspring by virgin<br />

birth or by vegetative reproduction; that is, reproduction without<br />

sexual fertilization of eggs.<br />

assortative mating Tendency of like to mate with like. It can<br />

be for a certain genotype (e.g., individuals with genotype AA<br />

tend to mate with other individuals of genotype AA) or phenotype<br />

(e.g., tall individuals mate with other tall individuals).<br />

atomistic (as applied to theory of inheritance) Inheritance in<br />

which the entities controlling heredity are relatively distinct,<br />

permanent, and capable of independent action. Mendelian<br />

inheritance is an atomistic theory because, in it, inheritance is<br />

controlled by distinct genes.<br />

autosome Any chromosome other than a sex chromosome.<br />

base The DNA is a chain of nucleotide units, and each unit consists<br />

of a backbone made of a sugar and a phosphate group, with<br />

a nitrogenous base attached. The base in a unit is one of adenine<br />

(A), guanine (G), cytosine (C), or thymine (T). In RNA, uracil<br />

(U) is used instead of thymine. A and G belong to the chemical<br />

class called purines; C, T, and U are pyrimidines.<br />

Batesian mimicry A kind of mimicry in which one nonpoisonous<br />

species (the Batesian mimic) mimics another poisonous<br />

species.<br />

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