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Evolution__3rd_Edition

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viii Full Contents<br />

3.5 Ring “species” show that the variation within a species can be<br />

extensive enough to produce a new species 50<br />

3.6 New, reproductively distinct species can be produced experimentally 53<br />

3.7 Small-scale observations can be extrapolated over the long term 54<br />

3.8 Groups of living things have homologous similarities 55<br />

3.9 Different homologies are correlated, and can be hierarchically classified 61<br />

3.10 Fossil evidence exists for the transformation of species 64<br />

3.11 The order of the main groups in the fossil record suggests they have<br />

evolutionary relationships 65<br />

3.12 Summary of the evidence for evolution 66<br />

3.13 Creationism offers no explanation of adaptation 67<br />

3.14 Modern “scientific creationism” is scientifically untenable 67<br />

Summary Further reading Study and review questions<br />

4. Natural Selection and Variation 71<br />

4.1 In nature, there is a struggle for existence 72<br />

4.2 Natural selection operates if some conditions are met 74<br />

4.3 Natural selection explains both evolution and adaptation 75<br />

4.4 Natural selection can be directional, stabilizing, or disruptive 76<br />

4.5 Variation in natural populations is widespread 81<br />

4.6 Organisms in a population vary in reproductive success 85<br />

4.7 New variation is generated by mutation and recombination 87<br />

4.8 Variation created by recombination and mutation is random<br />

with respect to the direction of adaptation 88<br />

Summary Further reading Study and review questions<br />

PART 2. EVOLUTIONARY GENETICS 93<br />

5. The Theory of Natural Selection 95<br />

5.1 Population genetics is concerned with genotype and gene frequencies 96<br />

5.2 An elementary population genetic model has four main steps 97<br />

5.3 Genotype frequencies in the absence of selection go to the Hardy–<br />

Weinberg equilibrium 98<br />

5.4 We can test, by simple observation, whether genotypes in a population<br />

are at the Hardy–Weinberg equilibrium 102<br />

5.5 The Hardy–Weinberg theorem is important conceptually,<br />

historically, in practical research, and in the workings of<br />

theoretical models 103<br />

5.6 The simplest model of selection is for one favored allele at one locus 104<br />

5.7 The model of selection can be applied to the peppered moth 108<br />

5.7.1 Industrial melanism in moths evolved by natural selection 108<br />

5.7.2 One estimate of the fitnesses is made using the rate of change<br />

in gene frequencies 109<br />

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