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Evolution__3rd_Edition

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22 PART 1 / Introduction<br />

DNA is carried in different ways in<br />

prokaryotic and in eukaryotic cells<br />

Figure 2.1<br />

The cells of a body have a fine<br />

structure (or “ultrastructure”)<br />

made up of a number of<br />

organelles. Not all the<br />

organelles illustrated here are<br />

found in all cells. Animal and<br />

fungal cells, for example,<br />

lack plastids; but all<br />

photosynthesizing organisms<br />

have them. Eukaryotes (i.e.,<br />

all plants and animals) have<br />

complex cells with a separate<br />

nucleus. Within the nucleus the<br />

DNA is here illustrated in the<br />

diffuse form called chromatin;<br />

when the cell divides, the<br />

chromatin coalesces into<br />

structures called chromosomes.<br />

Prokaryotes are simpler<br />

organisms, particularly<br />

bacteria, and they lack a distinct<br />

nucleus; their DNA lies naked<br />

within the cell.<br />

2.1 Inheritance is caused by DNA molecules, which are<br />

physically passed from parent to offspring<br />

The molecule called DNA (deoxyribose nucleic acid) provides the physical mechanism<br />

of heredity in almost all living creatures. The DNA carries the information used to<br />

build a new body, and to differentiate its various body parts. DNA molecules exist<br />

inside almost all the cells of a body, and in all the reproductive cells (or gametes). Its<br />

precise location in the cell depends on cell type.<br />

There are two main types of cell: eukaryotic and prokaryotic (Figure 2.1). Eukaryotic<br />

cells have a complex internal structure, including internal organelles and a distinct<br />

region, surrounded by a membrane, called the nucleus. Eukaryotic DNA exists within<br />

the nucleus. Prokaryotic cells are simpler and have no nucleus. Prokaryotic DNA lies<br />

within the cell, but in no particular region. All complex multicellular organisms,<br />

including all plants and animals, are built of eukaryotic cells. Fungi are also eukaryotic;<br />

some fungi are multicellular (such as mushrooms) others are unicellular (such as<br />

baker’s and brewer’s yeast, Saccharomyces cerevisiae). Protozoans, most of which (such<br />

as amebas) are unicellular, are the other main group of eukaryotes. Bacteria and<br />

Archaea are the two kinds of life in which the cells are prokaryotic.<br />

Within a eukaryotic cell nucleus, the DNA is physically carried in structures called<br />

chromosomes. Chromosomes can be seen through a light microscope at certain stages in<br />

the cell cycle. Individuals of different species characteristically have different numbers<br />

of chromosomes a each individual human has 46, for example, whereas a fruitfly<br />

Drosophila melanogaster has eight, and other species have other numbers. The finer<br />

structure of the DNA is too small to be seen directly, but it can be inferred by the<br />

method of X-ray diffraction. The molecular structure of DNA was worked out by<br />

Watson and Crick in 1953.<br />

Non-cellulosic cell wall<br />

Cell membrane<br />

Nucleoid<br />

Small ribosomes<br />

Flagellum<br />

Prokaryote<br />

Golgi body<br />

Large<br />

ribosome<br />

Endoplasmic<br />

reticulum<br />

Chromatin<br />

Eukaryote<br />

Mitochondria Plastid<br />

Kinetosome (9 + 0)<br />

Kinetochores<br />

Thylakoids<br />

Plastid inner membrane<br />

Plastid outer membrane<br />

Nucleus<br />

Nucleolus<br />

Nuclear membrane<br />

Cell membrane<br />

Cell wall (cellulose)<br />

Cilia (9 + 2)<br />

Cell membrane<br />

..

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