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Walia Special Edition on the Bale Mountains (2011) - Zoologische ...

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History and Physical Features of <strong>the</strong> Ethiopian Highlands<br />

The Ethiopian highlands were formed during <strong>the</strong> Oligocene and Miocene geological periods, between<br />

38 and 7 milli<strong>on</strong> years ago (Miehe and Miehe 1994), before <strong>the</strong> formati<strong>on</strong> of <strong>the</strong> Rift Valley that<br />

splits <strong>the</strong> highlands into two (Yalden and Largen 1992). The highlands have a lower limit of about<br />

1100 m a.s.l. and an area of 519 278 km2 (Williams et al. 2004). Ethiopia c<strong>on</strong>tains more than 80% of<br />

<strong>the</strong> highland massif above 3000m a.s.l. <strong>on</strong> <strong>the</strong> African c<strong>on</strong>tinent (Yalden 1983). The highlands (land<br />

above 1500 m a.s.l.) c<strong>on</strong>stitute around 43% of <strong>the</strong> country and are <strong>the</strong> most densely populated areas<br />

in <strong>the</strong> country, inhabited by 88% of Ethiopia’s human and 65% of its livestock populati<strong>on</strong> (Yalden<br />

1983; CSE 1997; EWNHS 2001). This has led to land degradati<strong>on</strong> through agriculture, overgrazing<br />

and soil erosi<strong>on</strong>, which are <strong>the</strong> most serious threats to <strong>the</strong> sustainability of <strong>the</strong>se biologically and<br />

ecologically important ecosystems (EWNHS 1996; Stephens et al. 2001; Williams et al. 2004).<br />

The <strong>Bale</strong> <strong>Mountains</strong> are situated in <strong>the</strong> sou<strong>the</strong>ast highlands of Ethiopia, geographically<br />

separated from <strong>the</strong> western and central highlands of <strong>the</strong> country by <strong>the</strong> Rift Valley. Located in <strong>the</strong><br />

centre of <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong>, <strong>the</strong> BMNP encompasses an area of 2200 km2 . The Nati<strong>on</strong>al Park<br />

c<strong>on</strong>tains a landscape ranging from 1500 m a.s.l. to 4377 m a.s.l. Five main vegetati<strong>on</strong> z<strong>on</strong>es are<br />

designated in this Nati<strong>on</strong>al Park: <strong>the</strong> nor<strong>the</strong>rn Grasslands (Gaysay valley), <strong>the</strong> nor<strong>the</strong>rn woodlands,<br />

ericaceous forest, <strong>the</strong> Afroalpine moorland and grassland and <strong>the</strong> Harenna Forest (Hillman 1986;<br />

Miehe and Miehe 1994; Asefa 2006-07).<br />

The BMNP area experiences two rainy seas<strong>on</strong>s: heavy rain and small rain. The heavy rain<br />

is from July to October, with <strong>the</strong> highest peak in <strong>the</strong> August and <strong>the</strong> small rain from March to June,<br />

with <strong>the</strong> highest peak in <strong>the</strong> April. The highest temperature recorded is 18.4oC in February and <strong>the</strong><br />

lowest is 1.4oC in January (Hillman 1986; Refera and Bekele 2002).<br />

Provisi<strong>on</strong>al Checklist and Endemic Species of <strong>the</strong> BMNP<br />

In this review paper, nomenclature mostly follows <strong>the</strong> names used by Hillman (1993). However,<br />

<strong>the</strong> materials referred by Hillman (1986), Yalden (1988) and Hillman (1993) to Crocidura fumosa<br />

from <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> were found to be a mis-identificati<strong>on</strong> (Yalden and Largen 1992); thus,<br />

as used in this paper, <strong>the</strong> <strong>on</strong>e from <strong>the</strong> Harenna forest in <strong>the</strong> south side of those mountains is C.<br />

thalia, whereas that from <strong>the</strong> north side is C. glassi. Also, <strong>the</strong> material <strong>the</strong>se authors referred to as<br />

C. baileyi is in fact C. lucina (Yalden and Largen 1992). For <strong>the</strong> remainder of species, which were<br />

not menti<strong>on</strong>ed in Hillman’s (1993) list, names follow Kingd<strong>on</strong> (2004). Deviati<strong>on</strong>s from <strong>the</strong>se, when<br />

deemed necessary, are indicated where appropriate.<br />

In his pi<strong>on</strong>eer work of attempting to compile a checklist of mammals of <strong>the</strong> BMNP, Hillman<br />

(1986) listed <strong>the</strong> occurrence of 47 mammal species in <strong>the</strong> area. Six years later, he produced a revised<br />

checklist in which he listed 66 species (Hillman 1993). Thereafter, an updated checklist of mammals<br />

in <strong>the</strong> Nati<strong>on</strong>al Park has not been published. However, unpublished documents <strong>on</strong> mammals of <strong>the</strong><br />

BMNP suggest a total of 77 species in <strong>the</strong> park (Williams 2002; BMNP 2003).<br />

<str<strong>on</strong>g>Walia</str<strong>on</strong>g>-<str<strong>on</strong>g>Special</str<strong>on</strong>g> <str<strong>on</strong>g>Editi<strong>on</strong></str<strong>on</strong>g> <strong>on</strong> <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> 5

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