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Walia Special Edition on the Bale Mountains (2011) - Zoologische ...

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forest, and this indicates that <strong>the</strong>y are affected by oligotrophic waters. The difference in pH and<br />

nutrient status between glades and forest decreases with decreasing altitude. The water originating<br />

from high altitude is naturally oligotrophic and it progressively charges with cati<strong>on</strong>s downstream.<br />

Al<strong>on</strong>gside this simple model, we must c<strong>on</strong>sider in this case o<strong>the</strong>r mechanisms resulting in water<br />

oligotrophy, such as podzolizati<strong>on</strong> and drainage from acidic badrocks.<br />

In circumstance where SOM turnover rates are reduced, such as anaerobic c<strong>on</strong>diti<strong>on</strong> due to<br />

water logging, low temperature due to high altitude, extremely acidic parent materials or producti<strong>on</strong><br />

of low quality OM (such as litter from c<strong>on</strong>iferous trees, Erica spp., etc.), <strong>the</strong> net accumulati<strong>on</strong> of<br />

organic acids will lead to podzolizati<strong>on</strong> even in <strong>the</strong> tropics (FAO 2001). Hydromorphic podzols are<br />

caused by waterlogging and have predominant lateral spread, with groundwater enriched in OM<br />

travelling l<strong>on</strong>g distances (FAO 2001) and eventually emerges at lower altitudes or in depressi<strong>on</strong>s<br />

in <strong>the</strong> form of black rivers (such as <strong>the</strong> Rio Negro in S. America) and lakes. pH levels are a good<br />

indicati<strong>on</strong> of a cause for podzolizati<strong>on</strong>, favoured at pH5<br />

(Shoji et al. 1993).<br />

In <strong>the</strong> toposequence studied, looking at pH values, we could predict that podzolizati<strong>on</strong> would<br />

occur above S-1850 and ferralitizati<strong>on</strong> downstream. Evidence of ferralitizati<strong>on</strong> is found in <strong>the</strong> high<br />

chroma of <strong>the</strong> soil in <strong>the</strong> landscape surrounding Delo-Mena (South of <strong>the</strong> studied toposequence),<br />

indicating enrichment of sesquioxides in <strong>the</strong> topsoil. In accordance to what described by Chauvel<br />

et al. (1996) for <strong>the</strong> Rio Negro basin, if podzolizati<strong>on</strong> occurs at higher altitudes, it could generate a<br />

deep flow of organic acids and Al that would extend into <strong>the</strong> ferralitic z<strong>on</strong>e downstream; this flow<br />

will stop in <strong>the</strong> soil at saturati<strong>on</strong> or will emerge in depressi<strong>on</strong>s (black lakes) or streams (black rivers)<br />

at lower altitudes (Fig. 3).<br />

Figure 3. Sketch diagram (not in scale) summarizing <strong>the</strong> elements supporting <strong>the</strong> <strong>the</strong>ory of a deep<br />

“black water” flow, and integrating <strong>the</strong>m in a simplified model of <strong>the</strong> escarpment.<br />

<str<strong>on</strong>g>Walia</str<strong>on</strong>g>-<str<strong>on</strong>g>Special</str<strong>on</strong>g> <str<strong>on</strong>g>Editi<strong>on</strong></str<strong>on</strong>g> <strong>on</strong> <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> 140

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