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Walia Special Edition on the Bale Mountains (2011) - Zoologische ...

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Ericaceous vegetati<strong>on</strong>. The present study attempts to provide a descripti<strong>on</strong> of plant communities and<br />

an analysis of <strong>the</strong> structure of woody species in Ericaceous vegetati<strong>on</strong> al<strong>on</strong>g <strong>the</strong> sou<strong>the</strong>rn slopes of<br />

<strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong>.<br />

Descripti<strong>on</strong> of <strong>the</strong> Study Area<br />

Floristic background<br />

The botanical explorati<strong>on</strong> of <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> started in 1906 by Neuman and Ellenbeck (cited in<br />

Miehe and Miehe 1994), followed by Smeds (1959), and Mo<strong>on</strong>ey (1963). In a series of publicati<strong>on</strong>s<br />

Hedberg (1951, 1964, 1975, 1986) made important analyses of <strong>the</strong> vegetati<strong>on</strong> and ecology of<br />

Afroalpine regi<strong>on</strong>s in Ethiopia. Weinert (1981), Weinert and Mazurek (1984) and Uhlig (1988)<br />

also published ecological studies <strong>on</strong> <strong>the</strong> vegetati<strong>on</strong> of <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong>. Research <strong>on</strong> <strong>the</strong> floristic<br />

compositi<strong>on</strong> and physiognomy of <strong>the</strong> m<strong>on</strong>tane vegetati<strong>on</strong> was c<strong>on</strong>ducted by Mesfin Tadesse (1986),<br />

Friis (1986), Lisanework Negatu and Mesfin Tadesse (1989), and Uhlig (1988, 1991). However,<br />

except for Hedberg (1986), Menassie Gashaw and Masresha Fetene (1996), and Miehe and Miehe<br />

(1994), most of <strong>the</strong>se studies were c<strong>on</strong>fined to <strong>the</strong> m<strong>on</strong>tane forest belt.<br />

Based <strong>on</strong> its physiognomy, Friis (1986) recognized <strong>the</strong> z<strong>on</strong>e between 3200 m and 3500 m<br />

as <strong>the</strong> uppermost forest z<strong>on</strong>e. This vegetati<strong>on</strong> z<strong>on</strong>e is relatively little disturbed by humans (Miehe<br />

and Miehe 1994). The main c<strong>on</strong>stituent of this z<strong>on</strong>e, Erica trimera, forms trees up to 15 m in<br />

height. The vegetati<strong>on</strong> differs c<strong>on</strong>siderably in physiognomy from that of higher elevati<strong>on</strong>s. Friis<br />

(1986) recognizes fire as <strong>the</strong> primary reas<strong>on</strong> for this physiognomic change, and altitude as a possible<br />

additi<strong>on</strong>al factor.<br />

More detailed data <strong>on</strong> <strong>the</strong> Ericaceous vegetati<strong>on</strong> of <strong>Bale</strong> were provided by Miehe and Miehe<br />

(1994), who compared and c<strong>on</strong>trasted <strong>the</strong>ir findings globally and regi<strong>on</strong>ally. The giant hea<strong>the</strong>rs <strong>on</strong><br />

<strong>the</strong> sou<strong>the</strong>rn slopes of <strong>the</strong> mountains harbor epiphytic communities that are crucial in buffering<br />

water deifiet between m<strong>on</strong>tane and Afroalpine vegetati<strong>on</strong>. These epiphytic mosses gradually release<br />

<strong>the</strong> absorbed moisture; it was estimated that up to 50,000 litters per ha are absorbed in ano<strong>the</strong>r east<br />

African cloud forest (Pocs 1976). The cryptograms also serve as bio-indicators of envir<strong>on</strong>mental<br />

c<strong>on</strong>diti<strong>on</strong>s.<br />

Geology and climate<br />

The study area is located at <strong>the</strong> sou<strong>the</strong>rn slope of <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong>, at <strong>the</strong> Harenna escarpment<br />

between 6°45′ and 7°N and 39°45′ and 39°40′ E. The rocks are of volcanic origin. They are<br />

predominantly trachytes but also include rhyolites, basalts and associated agglomerates and tuffs<br />

(Weinert and Mazurek 1984). Although detailed informati<strong>on</strong> about glaciati<strong>on</strong>s is still lacking, <strong>the</strong><br />

current land forms in <strong>the</strong> mountains appear to have resulted from tect<strong>on</strong>ic activity and glaciati<strong>on</strong>s. At<br />

least two glacial periods (18,000 BP and 2,000 BP) are documented from <strong>the</strong> mountains (B<strong>on</strong>nefille<br />

1993).<br />

<str<strong>on</strong>g>Walia</str<strong>on</strong>g>-<str<strong>on</strong>g>Special</str<strong>on</strong>g> <str<strong>on</strong>g>Editi<strong>on</strong></str<strong>on</strong>g> <strong>on</strong> <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> 160

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