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Walia Special Edition on the Bale Mountains (2011) - Zoologische ...

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interacti<strong>on</strong>s? 2) Would such classificati<strong>on</strong> of species in to communities be reliably detectable in <strong>the</strong><br />

field? Fur<strong>the</strong>r questi<strong>on</strong>s are as follows: Can <strong>on</strong>e c<strong>on</strong>sider <strong>the</strong> whole bird assemblage of <strong>the</strong> BMNP<br />

as members of a single community or several communities? If <strong>the</strong> latter, what is <strong>the</strong> species pool<br />

of each community? Is <strong>the</strong>re intercommunity difference in compositi<strong>on</strong> across habitat types? Can<br />

such classificati<strong>on</strong> reflect <strong>the</strong> distributi<strong>on</strong> of resident birds across different habitat types within each<br />

ecosystem?<br />

Methods<br />

Study area<br />

The <strong>Bale</strong> <strong>Mountains</strong> Nati<strong>on</strong>al Park located at 6030’- 7o00’ N, 39030’- 39055 E is part of <strong>the</strong> South-<br />

Eastern highland massif in Ethiopia and c<strong>on</strong>tains <strong>the</strong> largest extent of Afro-alpine habitat in <strong>the</strong><br />

c<strong>on</strong>tinent (Sillero-Zubiri 1995). It encompasses an area of 2200 km2 with elevati<strong>on</strong> varying between<br />

1400 m to 4400 m a.s.l. (Hillman 1986). Topography within <strong>the</strong> BMNP can be divided in to three<br />

categories by elevati<strong>on</strong>-<strong>the</strong> nor<strong>the</strong>rn slopes between 3000 and 3800 m a.s.l.; <strong>the</strong> central plateau<br />

and peaks between 3800 and 4400 m a.s.l.; and <strong>the</strong> sou<strong>the</strong>rn escarpment from 1400-3800 m a.s.l.<br />

(Hillman 1988). Depending <strong>on</strong> slope, aspect and soil types, each of <strong>the</strong>se areas in <strong>the</strong> park has its<br />

own characteristic vegetati<strong>on</strong>. In <strong>the</strong> nor<strong>the</strong>rn slopes a mix of Juniperus/Hagenia/Hypericum where<br />

m<strong>on</strong>tane grassland and heath dominates near <strong>the</strong> tree line; sparse afro-alpine vegetati<strong>on</strong> at its center<br />

and; and <strong>the</strong> vegetati<strong>on</strong> in <strong>the</strong> sou<strong>the</strong>rn escarpment c<strong>on</strong>stitutes Erica/Hypericum/Hagenia/Aningeria<br />

and Podocarpus. A bimodal local climate with two wet seas<strong>on</strong>s that have heavy and small rains<br />

is characteristic to <strong>the</strong> site. The heavy rains occur from July to October, with <strong>the</strong> highest peak in<br />

August and <strong>the</strong> small rains from March to June, with a peak in April. Records show that this area<br />

experiences temperature extremities particularly in areas of <strong>the</strong> highest altitudes during <strong>the</strong> dry<br />

seas<strong>on</strong> and more or less <strong>the</strong> same pattern of warm during <strong>the</strong> wet seas<strong>on</strong>. The highest temperature is<br />

18.4 Co in February and <strong>the</strong> lowest is 1.4 o in January (Hillman 1986).<br />

Structuring of <strong>the</strong> avi-fauna of <strong>the</strong> BMNP<br />

Censuses c<strong>on</strong>ducted between November 2002 and October 2005 determined a total list of resident<br />

birds composed of 117 species in <strong>the</strong> Nati<strong>on</strong>al Park (NP). The range of species includes wetland<br />

birds to small passerines that inhabit forest and afro-alpine moorland habitats. As much as <strong>the</strong><br />

habitat preference of groups of species, <strong>the</strong>ir diet and related ecological and morphological features<br />

are diverse (Fry et al. 1992; Zimmerman et al. 1996; Sincliar and Ryan 2003). This suggests <strong>the</strong><br />

total bird assemblage of <strong>the</strong> NP may c<strong>on</strong>tain a group of species that may be assembled to form more<br />

than <strong>on</strong>e community. The possibility of such phenomena was explored by gleaning data from <strong>the</strong><br />

literature that helped to evaluate <strong>the</strong> potential of interspecifc interacti<strong>on</strong>s.<br />

Interacti<strong>on</strong> am<strong>on</strong>gst species is defined as <strong>the</strong> ability of each to affect <strong>the</strong> resource level<br />

that is available to <strong>on</strong>e or more of species that co-occur with it both in space and time (Daim<strong>on</strong>d<br />

1975; Rice and Kr<strong>on</strong>lund 1997). Resources for a species include food, breeding habitat and cover<br />

<str<strong>on</strong>g>Walia</str<strong>on</strong>g>-<str<strong>on</strong>g>Special</str<strong>on</strong>g> <str<strong>on</strong>g>Editi<strong>on</strong></str<strong>on</strong>g> <strong>on</strong> <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> 17

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