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Walia Special Edition on the Bale Mountains (2011) - Zoologische ...

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In c<strong>on</strong>trast to <strong>the</strong> nor<strong>the</strong>rn highlands, sou<strong>the</strong>rn Ethiopia is within <strong>the</strong> East African climatic<br />

domain, which is influenced during <strong>the</strong> larger part of <strong>the</strong> year by south-easterlies originating over<br />

<strong>the</strong> Indian Ocean. As in most Ethiopian highlands, <strong>the</strong> inter-tropical c<strong>on</strong>vergence z<strong>on</strong>e (ITCZ),<br />

plus altitudinal and topographic influences all affect <strong>the</strong> distributi<strong>on</strong> of <strong>the</strong> precipitati<strong>on</strong> in <strong>the</strong> <strong>Bale</strong><br />

<strong>Mountains</strong>. Annual rainfall ranges between 600-1500 (2000) mm depending <strong>on</strong> <strong>the</strong> relief and altitude<br />

Diurnal variability in temperature in <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> is higher than its seas<strong>on</strong>al variati<strong>on</strong>.<br />

A minimum temperature of -15°C was recorded by Hillman (1986) <strong>on</strong> <strong>the</strong> Plateau (3850 m), while<br />

Miehe and Miehe (1994) recorded a night-time minimum temperature of -3°C in <strong>the</strong> sparsely<br />

vegetated areas of <strong>the</strong> Ericaceous belt. Soliflucti<strong>on</strong> is comm<strong>on</strong> in <strong>the</strong> Afroalpine belt and in <strong>the</strong><br />

upper parts of <strong>the</strong> Ericaceous vegetati<strong>on</strong>.<br />

There is evidence of early settlements in some valleys and plains. Thus, <strong>the</strong> area has probably<br />

been populated since ancient times, but human land use and barley producti<strong>on</strong> in <strong>the</strong> Ericaceous and<br />

Afroalpine belt has increased tremendously with <strong>the</strong> c<strong>on</strong>structi<strong>on</strong> of an-all-wea<strong>the</strong>r road traversing<br />

<strong>the</strong> plateau. However, <strong>the</strong> highlands of <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> are still less densely populated than<br />

<strong>the</strong> Semien <strong>Mountains</strong> in northwestern Ethiopia (populati<strong>on</strong> densities are 23.4/km2 and 52.6/km2 ,<br />

respectively). Barley is cultivated in <strong>Bale</strong> 600-800 m lower than in Semien, and this is due to <strong>the</strong><br />

transhumant mode of living in <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong>.<br />

Vegetati<strong>on</strong> Sampling<br />

The current study covered vegetati<strong>on</strong> in <strong>the</strong> Ericaceous belt of <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> al<strong>on</strong>g an altitudinal<br />

gradient ranging from 3000 to 4200 m. Transects were established in homogeneous patches of<br />

vegetati<strong>on</strong> (Mueller-Dombois and Ellenberg 1974). Plots of 15 x 15 m were sampled at intervals<br />

of 20 m lateral distance between <strong>the</strong> plots. For each plot, <strong>on</strong>e sub-plot of 2 x 2 m was surveyed<br />

for <strong>the</strong> herbaceous vegetati<strong>on</strong>. All vascular plants in each plot were recorded by estimating <strong>the</strong>ir<br />

cover-abundances using <strong>the</strong> 9-level scale of Braun Blanquet as modified by Van der Maarel (1979).<br />

The size class structure of all tree and shrub species was also assessed in each plot by recording<br />

<strong>the</strong> height and diameter at breast/stump height (DBH/DSH) of all stems > 2.5 cm were taken. For<br />

multi-stemmed trees, each stem was recorded as individual tree. Species were identified <strong>on</strong> site with<br />

<strong>the</strong> help of trained tax<strong>on</strong>omist and specimens s were collected for identificati<strong>on</strong> for using flora of<br />

Ethiopia. Seedlings (below 1.5 m) and saplings (above 1.5 meter but below 2 meter) of juvenile<br />

woody species were counted in a sub plot of 5 x 5 m in each quadrat; <strong>the</strong> total sample was thus 110<br />

plots and 110 sub plots.<br />

Data Analysis<br />

Vegetati<strong>on</strong> data were analyzed with <strong>the</strong> software package SYN-TAx 2000, using <strong>the</strong> hierarchical<br />

clustering by distance optimizati<strong>on</strong> (Podani 2000). The Soerensen similarity index was used as <strong>the</strong><br />

distance measure and <strong>the</strong> average linkage as <strong>the</strong> cluster algorithm. Species diversity was measured<br />

using <strong>the</strong> Shann<strong>on</strong>-Weaver index of diversity (Krebs 1989). As a proxy for <strong>the</strong> age structure of<br />

<str<strong>on</strong>g>Walia</str<strong>on</strong>g>-<str<strong>on</strong>g>Special</str<strong>on</strong>g> <str<strong>on</strong>g>Editi<strong>on</strong></str<strong>on</strong>g> <strong>on</strong> <strong>the</strong> <strong>Bale</strong> <strong>Mountains</strong> 161

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