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244<br />

INSECTA, COLEOPTERA.<br />

South and Central America east of <strong>the</strong> Andes and north of <strong>the</strong> River Plate,<br />

and furnishes a large share <strong>in</strong> <strong>the</strong> constitution of North America, receiv<strong>in</strong>g <strong>in</strong><br />

turn a very perceptible t<strong>in</strong>ge from <strong>the</strong> microtypal stirps. The microtypal<br />

stirps <strong>in</strong>cludes Europe, Asia north of <strong>the</strong> Himalayas, Eastern North America<br />

(so far as not modified by <strong>the</strong> Brazilian element), and (<strong>in</strong> a less degree) Northwest<br />

America, California, part of Mexico, Peru, Chili, <strong>the</strong> Argent<strong>in</strong>e Republic<br />

south of Tucuman, Patagonia, Tierra del Fuego, Polynesia, New Zealand,<br />

and Australia. The apparent paradox of <strong>in</strong>clud<strong>in</strong>g <strong>the</strong> first and last of<br />

<strong>the</strong>se countries <strong>in</strong> <strong>the</strong> same stirps is endeavoured to be accounted for by <strong>the</strong><br />

<strong>in</strong>ference, from <strong>the</strong> fact of <strong>the</strong> Eocene flora of Europe hav<strong>in</strong>g many po<strong>in</strong>ts of<br />

correlation with <strong>the</strong> present Australian flora, and from <strong>the</strong> author's supposition<br />

of <strong>the</strong> probability of <strong>the</strong> European Eocene beetle-fauna hav<strong>in</strong>g been <strong>the</strong> same<br />

as <strong>the</strong> Miocene (based upon Heer's proof that <strong>the</strong> Miocene beetle-fauna of<br />

Europe was of <strong>the</strong> same type as at present), that <strong>the</strong> Eocene fauna of Em-ope<br />

has (like <strong>the</strong> Eocene flora of Australia) survived <strong>in</strong> its Eocene form down to<br />

<strong>the</strong> present day. The author's conviction is that <strong>the</strong>re have been almost certa<strong>in</strong>ly<br />

two great cont<strong>in</strong>ental routes of communication between <strong>the</strong> nor<strong>the</strong>rn<br />

and sou<strong>the</strong>rn hemispheres—one now at <strong>the</strong> bottom of <strong>the</strong> Pacific, <strong>the</strong> o<strong>the</strong>r<br />

of <strong>the</strong> Atlantic. The beetle-fauna? of <strong>the</strong> Madeiran Isles and of <strong>the</strong> Azores, as<br />

enumerated by Wollaston and Crotch, are considered substantially to corroborate<br />

<strong>the</strong> microtypal stirps, <strong>the</strong> European element (hi<strong>the</strong>rto held to have<br />

been <strong>in</strong>troduced) be<strong>in</strong>g treated as natural denizens. The entire fauna of St.<br />

Helena is discussed at some length (<strong>the</strong> -author admitt<strong>in</strong>g that that island is<br />

<strong>the</strong> crucial test of his hypo<strong>the</strong>sis of a communication between <strong>the</strong> nor<strong>the</strong>rn<br />

and sou<strong>the</strong>rn hemispheres by an Atlantic cont<strong>in</strong>ent), and is regarded as belong<strong>in</strong>g<br />

to <strong>the</strong> Atlantic subfauna of <strong>the</strong> microtypal stirps. Eastward, this<br />

stirps is traced through Siberia, Ch<strong>in</strong>a (due stress be<strong>in</strong>g given to <strong>the</strong> prevalence<br />

<strong>the</strong>re of European forms), and Japan (where, however, as <strong>in</strong> Ch<strong>in</strong>a, <strong>the</strong><br />

Hymenoptera are wholly different from <strong>the</strong> European type), to North America,<br />

<strong>the</strong>nce, vid California, to Mexico and <strong>the</strong> o<strong>the</strong>r South-American countries<br />

above mentioned. Coast- and cave-beetles, and <strong>the</strong> occasional occurrence<br />

of European genera, are relied upon as strong evidence of all <strong>the</strong>se belong<strong>in</strong>g<br />

to <strong>the</strong> same stirps,—<strong>the</strong> occurrence of South-African forms <strong>in</strong> South<br />

America be<strong>in</strong>g accounted for by a submerged Patagonian cont<strong>in</strong>ent, which<br />

may have existed at three different epochs. The non-coral isles of Polynesia<br />

are claimed as microtypal, aga<strong>in</strong> chiefly on <strong>the</strong> ground of <strong>the</strong>ir possess<strong>in</strong>g<br />

British forms ; and <strong>the</strong> European aff<strong>in</strong>ities of a very considerable portion of<br />

<strong>the</strong> <strong>in</strong>gredients of <strong>the</strong> Coleopterous fauna of Australia are stated to be without<br />

doubt,—some peculiarly Australian forms be<strong>in</strong>g also thought to be local<br />

representatives of South-American (and <strong>the</strong>refore) European races. The<br />

common want of certa<strong>in</strong> conspicuous families or genera by Australia and <strong>the</strong><br />

microtypal stirps <strong>in</strong> o<strong>the</strong>r countries is relied upon by <strong>the</strong> author as a<br />

connect<strong>in</strong>g l<strong>in</strong>k; and <strong>the</strong> Miocene epoch supplies some miss<strong>in</strong>g forms.<br />

As regards <strong>the</strong> Indo-African stirps, <strong>the</strong> author's chief po<strong>in</strong>ts are his present<br />

conviction (opposed to Wallace's op<strong>in</strong>ion) that <strong>the</strong> Coleoptera of <strong>the</strong> New-<br />

Gu<strong>in</strong>ea islands are essentially Indo-Malayan,—that, although Africa (south of<br />

<strong>the</strong> Sahara) is better entitled to claim rank as a separate prov<strong>in</strong>ce than India,<br />

<strong>the</strong>re was but one orig<strong>in</strong>al stock for both, as most of <strong>the</strong> genera occurr<strong>in</strong>g <strong>in</strong><br />

one are found, under suitable conditions, <strong>in</strong> <strong>the</strong> o<strong>the</strong>r, and some large groups<br />

(e. g. Staphyliuidce) are absent <strong>in</strong> both of <strong>the</strong>m,—that Africa was disjo<strong>in</strong>ed

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