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Series editors' preface - Wood Tools

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about the same size as the width of a cotton<br />

fibre, or of a very large bacterial cell.<br />

Fungi have two basic kinds of structures,<br />

specialized for growth and for reproduction.<br />

The vegetative structure responsible for<br />

growth is the threadlike filament called the<br />

hypha. Collectively, hyphae are referred to as<br />

the mycelium. The diameter of the hyphae<br />

varies considerably but mostly falls in the<br />

range 2–20 μm. The principal reproductive<br />

unit is the spore. The relationship between the<br />

spores and the hyphae varies. In the simplest<br />

cases, the spore is formed from cells of the<br />

hyphae that separate at maturity. In other<br />

cases, more complex relationships exist.<br />

Spores may arise for example from specialized<br />

stalks or branches borne by the hyphae rather<br />

than directly from the hyphae themselves.<br />

Spores may be of two types. Under ideal<br />

growing conditions a type of spore may be<br />

formed rapidly and in large numbers that is<br />

suitable for rapid multiplication but has little<br />

resistance to adverse conditions. Another type<br />

of spore, suited to tiding the organism over<br />

periods unfavourable for growth, is produced<br />

under appropriate conditions. These may live<br />

for many years and can become widely distributed.<br />

Some of the fungi that cause deterioration<br />

reproduce by sexual methods while<br />

others do so asexually.<br />

Control of fungi Factors that influence the<br />

growth of fungi include temperature, humidity,<br />

light, oxygen and the food source. The<br />

temperature range inhabited by various different<br />

fungi goes from just above freezing to<br />

about 50 °C. For each species there is a<br />

preferred temperature range at which growth<br />

rate is optimum, a temperature below which<br />

growth will not occur and a temperature<br />

above which growth will not occur. The<br />

optimum for many fungi lies in the range<br />

15–30 °C. Many fungi will survive continuous<br />

freezing for several months but are less tolerant<br />

of repeated freeze–thaw cycles. Moist heat<br />

kills them more readily than dry heat. The<br />

critical average moisture content of a material<br />

for fungal growth is 20%. The corresponding<br />

RH is unlikely to be much below 70%. In<br />

conditions much drier than this fungi will not<br />

grow. Most fungi require a supply a supply of<br />

oxygen for growth but will operate independently<br />

of lighting conditions, although they<br />

General review of environment and deterioration 267<br />

may be damaged or killed by UV radiation.<br />

Fungi require sources of carbon, hydrogen,<br />

oxygen, nitrogen, sulphur, potassium, magnesium<br />

and phosphorous for growth. They may<br />

also require traces of iron, zinc, copper,<br />

calcium and manganese. Many naturally occurring<br />

compounds may be used by fungi as a<br />

source of carbon and energy. Glucose,<br />

sucrose, animal and vegetable fats and their<br />

component acids and glycerine can generally<br />

be used and some, but not all, fungi are able<br />

to utilize cellulose. Nitrogen may be assimilated<br />

as nitrates, ammonium salts, amino acids<br />

or proteins. Since the absence of any of these<br />

requirements will prevent or greatly inhibit<br />

growth, methods for control are based on<br />

eliminating or modifying one or more of these<br />

conditions by keeping objects clean and under<br />

suitable conditions of RH. Alternatively,<br />

poison, as used in wood preservatives, offers<br />

an effective and reliable means of protection<br />

in high risk situations. Methods of controlling<br />

fungal problems are further discussed by<br />

Strang and Dawson in Canadian Conservation<br />

Institute Technical Bulletin No. 12.<br />

The chief products of fungal metabolism are<br />

carbon dioxide and water. However, other<br />

products are also formed that themselves have<br />

a deleterious effect on objects. These include<br />

various organic acids, alcohols and esters. The<br />

formation of coloured material is also<br />

frequently conspicuous.<br />

Insects<br />

All insects are members of the class Insecta<br />

belonging to the vast phylum Arthropoda, the<br />

most complex and specialized group of invertebrates<br />

in the animal kingdom. About one<br />

million different insect species have been<br />

recorded to date though no one really knows<br />

how many species there are on the earth<br />

today. Between them, insects will eat virtually<br />

anything organic, including many of the<br />

materials used in the construction of furniture<br />

and wooden objects. They may also cause<br />

incidental damage, for example by their<br />

excreta. This causes them to be regarded as<br />

pests and for man to seek to eliminate them<br />

at least from environments in which valuable<br />

furniture is being maintained. Insect damage<br />

needs to be understood in relation to the<br />

habits of insects, their life cycle, the nature<br />

and distribution of the collections and the

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