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in Sri Lanka - Ministry of Environment

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Present Status <strong>of</strong> Dry-zone Flora <strong>in</strong> <strong>Sri</strong> <strong>Lanka</strong>Anoma Perera,Department <strong>of</strong> Botany, University <strong>of</strong> Peradeniya, Peradeniya.A. Introduction to the ecosystemThe dry land <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong> which is classically described as the dry and <strong>in</strong>termediate zones <strong>of</strong>the country, extends over about 66% <strong>of</strong> the total is land area, but nearly 85% <strong>of</strong> the naturalvegetation-cover <strong>of</strong> the country exists with<strong>in</strong> this region (Legg and Jewell, 1995). Three majornatural terrestrial vegetation types are found <strong>in</strong> the dry land <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>; viz, dry forests, savannaand grasslands. Depend<strong>in</strong>g on the variability <strong>in</strong> their location, climate, edaphic characteristicsand biotic impacts, it appears that several sub-categories <strong>of</strong> the above vegetation types canbe identified.Dry forest ecosystems<strong>Sri</strong> <strong>Lanka</strong>n dry forests are a part <strong>of</strong> the tropical ra<strong>in</strong>forest biome. These exhibit characteristicfeatures <strong>of</strong> tropical semi-deciduous or tropical seasonal forests <strong>of</strong> the world vegetation typesdescribed by Longman and Jenik (1987), <strong>in</strong> Archibold, (1995). Tropical semi-deciduous forestsgrow <strong>in</strong> low moisture conditions (e.g. annual ra<strong>in</strong> fall < 1200 mm) and are ma<strong>in</strong>ly found <strong>in</strong>Bundala National Park (Figure 1), southern part <strong>of</strong> Yala National Park and north-eastern part<strong>of</strong> Wilpaththu National Park. Presence <strong>of</strong> deciduous or leaf exchang<strong>in</strong>g species <strong>in</strong> the forestcanopy is a salient feature <strong>of</strong> these forests.In contrast, tropical seasonal forests (Figure 2) occur <strong>in</strong> areas where comparatively higherra<strong>in</strong>fall (mean annual ra<strong>in</strong>fall <strong>of</strong> 1200-1500 mm) is received and the major portion <strong>of</strong> the dryforests <strong>of</strong> the country falls <strong>in</strong>to this forest category. These forests grow taller than tropical semideciduousforests and show a clear stratification. Moreover, some evergreen species may bepresent <strong>in</strong> the canopy. However, the deviations <strong>in</strong> recorded physical features such as soil typeand elevation (Alwis and Eriyagama, 1969) as well as the slope <strong>of</strong> the terra<strong>in</strong> appear to result<strong>in</strong> spatial heterogeneity <strong>in</strong> the soil moisture contents. This has resulted <strong>in</strong> the formation <strong>of</strong>different forest communities which deviated from the typical Manilkara hexandra-Chloroxylonswietenia-Drypetes sepiaria community <strong>of</strong> lowland tropical seasonal forests.It has been reported that most <strong>of</strong> the dry forests <strong>in</strong> the lowlands had grown after the destruction<strong>of</strong> hydraulic civilization <strong>in</strong> the area around 13 th century (Brohier, 1941; de Rosayro, 1961) andtherefore, are secondary <strong>in</strong> orig<strong>in</strong>. Also, these forests are heavily disturbed by human activitiessuch as timber logg<strong>in</strong>g and frequent shift<strong>in</strong>g cultivation (de Jong et al., 2001; de Rosayro, 1961;Perera et al., 1995; Perera, 2001a) result<strong>in</strong>g <strong>in</strong> secondary forests or successional forests atdifferent ages (de Rosayro, 1961; Perera, 1998, 2001a). Natural regeneration <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>ndry forest species is reported to be very poor (de Rosayro, 1959; Holmes, 1957; Perera et al.,1995; Perera, 2001a; Rutnam, 1959) which is attributed to the poor quality and quantity <strong>of</strong> theseed ra<strong>in</strong> (Perera, 1998, 2004), soil seed bank (Perera, 1998a, 2000, 2004, 2005) and theseedl<strong>in</strong>g bank (Perera, 2001b, 2004). Repeated disturbances <strong>in</strong> secondary forests and lack <strong>of</strong>regeneration <strong>of</strong> high forest species may result <strong>in</strong> scrub jungle plagioclimaxes through deflectedsuccession but these are not a true climax vegetation type <strong>of</strong> the dry land <strong>of</strong> <strong>Sri</strong> <strong>Lanka</strong>. Such165

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