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Reduction and Elimination in Philosophy and the Sciences

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could mention <strong>the</strong> many sorts of isolat<strong>in</strong>g barriers (Section<br />

3) that prevent gene flow between dist<strong>in</strong>ct species (Wilson<br />

et al. forthcom<strong>in</strong>g). Despite its apparent plausibility, <strong>the</strong><br />

goal of this section is to present an objection aga<strong>in</strong>st HPC<br />

<strong>the</strong>ory.<br />

The properties <strong>in</strong> <strong>the</strong> cluster F can be ei<strong>the</strong>r <strong>in</strong>tr<strong>in</strong>sic<br />

or relational. Accord<strong>in</strong>g to <strong>the</strong> conceptual argument<br />

aga<strong>in</strong>st essentialism (Section 2), no <strong>in</strong>tr<strong>in</strong>sic property may<br />

count as necessary for membership <strong>in</strong> a species. But <strong>the</strong><br />

same argument is also effective aga<strong>in</strong>st Boyd’s f<strong>in</strong>ite<br />

disjunction of <strong>in</strong>tr<strong>in</strong>sic properties. 4 For if Okasha is right,<br />

<strong>the</strong>re is no boundary on <strong>the</strong> variation of <strong>in</strong>dividual traits<br />

among <strong>the</strong> members of a species. Thus, if <strong>the</strong> cluster F<br />

conta<strong>in</strong>s <strong>in</strong>tr<strong>in</strong>sic properties, <strong>the</strong>n given <strong>the</strong> rejection of<br />

essentialism F could not exclude any phenotypic or<br />

genotypic trait. If it did exclude such a trait, F would entail<br />

that not hav<strong>in</strong>g a certa<strong>in</strong> property is necessary for<br />

membership <strong>in</strong> a species. But this consequence cannot be<br />

right because F would <strong>the</strong>n not s<strong>in</strong>gle out <strong>in</strong>dividual<br />

species. To make this clear, consider an example.<br />

Suppose an unbounded disjunction of <strong>in</strong>tr<strong>in</strong>sic properties<br />

P1 v P2 v … <strong>and</strong> objects a <strong>and</strong> b both conta<strong>in</strong><strong>in</strong>g P1 <strong>and</strong> P2.<br />

Are <strong>the</strong>y both members of <strong>the</strong> same species? As <strong>the</strong><br />

disjunction is unbounded, <strong>the</strong> disjunction by itself cannot<br />

decide this question. Therefore, if we accept <strong>the</strong><br />

conceptual argument aga<strong>in</strong>st essentialism, F cannot<br />

conta<strong>in</strong> <strong>in</strong>tr<strong>in</strong>sic properties <strong>and</strong> so F can only conta<strong>in</strong><br />

relational properties.<br />

Now let us move to <strong>the</strong> second k<strong>in</strong>d of homeostasis<br />

(Homeostasis-II). As mentioned earlier, isolat<strong>in</strong>g barriers<br />

between species seem to count as Homeostasis-II. An<br />

important feature about isolat<strong>in</strong>g barriers is that <strong>the</strong>y are<br />

evolved characters between two species – e.g., courtship.<br />

S<strong>in</strong>ce isolat<strong>in</strong>g barriers are evolved characters, <strong>the</strong>y are<br />

analogous to phenotypic/genotypic traits: <strong>the</strong>y not only can<br />

vary through time, but also <strong>the</strong>re is no boundary to such<br />

variation. In contrast, non-<strong>in</strong>terbreed<strong>in</strong>g caused by<br />

geographic separation is not considered as an isolat<strong>in</strong>g<br />

barrier because it is not an evolved character (Ridley 2004:<br />

355).<br />

4 See Ereshefsky <strong>and</strong> Mat<strong>the</strong>n (2005), p. 9.<br />

Species, Variability, <strong>and</strong> Integration — Makmiller Pedroso<br />

But if <strong>the</strong> previous paragraph is correct, we can<br />

extend <strong>the</strong> argument used above aga<strong>in</strong>st Homeostasis-II.<br />

A f<strong>in</strong>ite disjunction of isolat<strong>in</strong>g barriers cannot s<strong>in</strong>gle out a<br />

species. O<strong>the</strong>rwise, we would have to accept that <strong>the</strong>re is<br />

some a priori impediment to how isolat<strong>in</strong>g barriers between<br />

two species can evolve. Hence, species cannot be<br />

dist<strong>in</strong>guished via Homeostasis-II.<br />

I have drawn two conclusions so far: (i) F can only<br />

conta<strong>in</strong> relational properties; <strong>and</strong> (ii) species cannot be<br />

dist<strong>in</strong>guished via Homeostasis-II. Because of (i) <strong>and</strong> (ii) it<br />

follows that, if species are HPC k<strong>in</strong>ds, <strong>the</strong>n <strong>the</strong>y are a<br />

cluster of relational properties where some of <strong>the</strong>se<br />

properties <strong>in</strong>duce <strong>the</strong> presence of o<strong>the</strong>rs (<strong>the</strong>reby, <strong>in</strong>tr<strong>in</strong>sic<br />

properties <strong>and</strong> Homeostasis-II are both excluded). If I am<br />

right about this, when applied to species, HPC <strong>the</strong>ory<br />

collapses <strong>in</strong>to a <strong>the</strong>ory that is no more explanatory than<br />

<strong>the</strong> species concepts <strong>the</strong>mselves. Boyd’s <strong>the</strong>ory can only<br />

state that some relational property such as <strong>in</strong>terbreed<strong>in</strong>g<br />

with con-specifics <strong>in</strong>duces o<strong>the</strong>r relational property like<br />

belong<strong>in</strong>g to <strong>the</strong> same gene pool as. So, although <strong>the</strong><br />

notions of homeostatic mechanisms or cluster of cooccurr<strong>in</strong>g<br />

properties seem to carry some additional<br />

explanatory power, I tried to show above that <strong>the</strong>se<br />

notions are irrelevant to comprehend<strong>in</strong>g what a species<br />

is.*<br />

Literature<br />

Boyd, R. 1991 “Realism, Anti-foundationalism, <strong>and</strong> <strong>the</strong> Enthusiasm<br />

for Natural K<strong>in</strong>ds” Philosophical Studies 61: 127-48.<br />

Boyd, R. 1999 “Homeostasis, Species, <strong>and</strong> Higher Taxa”, R. A.<br />

Wilson (ed.), Species: New Interdiscipl<strong>in</strong>ary Essays. Cambridge:<br />

MIT Press, 141-85.<br />

Ehrlich, P. <strong>and</strong> P. Raven (1969) “Differentiation of Populations”<br />

Science 165: 1228-32.<br />

Ereshefsky, M. 2001 The Poverty of L<strong>in</strong>naean Hierarchy Cambridge:<br />

Cambridge University Press.<br />

* Special thanks to Travis Dumsday for his helpful comments.<br />

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