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Benders'dictionary of nutrition and food technology

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92carbohydrate by difference It is relatively difficult to determinethe various carbohydrates present in <strong>food</strong>s, <strong>and</strong> an approximationis <strong>of</strong>ten made by subtracting the measured protein, fat, ash<strong>and</strong> water from the total weight. It is the sum <strong>of</strong> <strong>nutrition</strong>allyavailable carbohydrates (dextrins, starches <strong>and</strong> sugars); <strong>nutrition</strong>allyunavailable carbohydrate (pentosans, pectins, hemicelluloses<strong>and</strong> cellulose) <strong>and</strong> non-carbohydrates such as organicacids <strong>and</strong> lignins.carbohydrate loading Practice <strong>of</strong> some endurance athletes(e.g. marathon runners) in training for a major event; it consists<strong>of</strong> exercising to exhaustion, so depleting muscle glycogen, theneating a large carbohydrate-rich meal so as to replenish glycogenreserves with a higher than normal proportion <strong>of</strong> straightchain glycogen.carbohydrate metabolism See glucose metabolism.carbohydrate, unavailable A general term for those carbohydratespresent in <strong>food</strong>s that are not digested, <strong>and</strong> are thereforeexcluded from calculations <strong>of</strong> energy intake, although they maybe fermented by intestinal bacteria <strong>and</strong> yield some energy. Theterm includes both indigestible oligosaccharides <strong>and</strong> the variousnon-starch polysaccharides.See also fatty acids, volatile; starch, resistant.carbon dioxide, available See baking powder; flour, selfraising.carbon dioxide storage See packaging, modified atmosphere.g-carboxyglutamate A derivative <strong>of</strong> the amino acid glutamate(abbr Gla, M r 191.1) which is found in prothrombin <strong>and</strong> othercalcium-binding proteins involved in blood clotting. Its formationrequires vitamin k. Also occurs in the protein osteocalcin inbone, where it has a function in ensuring the correct crystallisation<strong>of</strong> bone mineral.carboxymethylcellulose See cellulose derivatives.carboxypeptidase E Enzyme (EC 3.4.17.10) that catalysescleavage <strong>of</strong> pro-insulin to insulin, <strong>and</strong> post-syntheticmodification <strong>of</strong> pro-opiomelanocortin <strong>and</strong> other peptidehormones.carboxypeptidases Enzymes (EC 3.4.17.1 <strong>and</strong> 2) secreted in thepancreatic juice that remove amino acids sequentially from thefree carboxyl end <strong>of</strong> a peptide or protein, i.e. exopeptidases.carcinogen Any compound that is capable <strong>of</strong> inducing cancer.carcinoid syndrome Condition in which there are metastases tothe liver <strong>of</strong> a carcinoid tumour <strong>of</strong> the enterochromaffin cells <strong>of</strong>the small intestine. The tumour produces a variety <strong>of</strong> physiologicallyactive amines, including histamine (which causes flushingreactions) <strong>and</strong> 5-hydroxytryptamine. The depletion <strong>of</strong> trypto-

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