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Benders'dictionary of nutrition and food technology

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282linoleic acid, conjugated Isomers <strong>of</strong> linoleic acid in which two ormore <strong>of</strong> the double bonds are conjugated (i.e. alternating withsingle bonds) rather than separated by a methylene bridge.a-linolenic acid An essential polyunsaturated fatty acid (C18:3ω3).g-linolenic acid A non-essential polyunsaturated fatty acid(C18:3 ω6), which has some pharmacological actions. Foundin oils from the seeds <strong>of</strong> evening primrose, borage <strong>and</strong>blackcurrant.linseed See flaxseed.liothyronine Obsolete name for the thyroid hormone triiodothyronine(T3).lipaemia Increase in blood lipids, as occurs normally after a meal.lipase Enzyme (EC 3.1.1.x) that hydrolyses triacylglycerols t<strong>of</strong>ree fatty acids <strong>and</strong> 2-mono-acylglycerol. Lipase secreted by thetongue <strong>and</strong> in gastric <strong>and</strong> pancreatic juice is EC 3.1.1.3; lipasesare also present in many seeds <strong>and</strong> grains. Final hydrolysis toyield glycerol is catalysed by acylglycerol lipase (EC 3.1.1.23).Most lipases have low specificity <strong>and</strong> will hydrolyse anytriacylglycerol. Sometimes responsible for the development <strong>of</strong>(hydrolytic) rancidity in stored <strong>food</strong>s, <strong>and</strong> the development <strong>of</strong>flavour in cheese.See also acid number; interesterification.lipase, clearing factor See lipase, lipoprotein.lipase, hormone-sensitive lipase in adipose tissue that is activatedin response to adrenaline, <strong>and</strong> inactivated in response toinsulin, so controlling release <strong>of</strong> free fatty acids as a metabolicfuel.lipase, lipoprotein lipase (EC 3.1.1.34) in muscle <strong>and</strong> adiposetissue that is responsible for the uptake <strong>of</strong> free fatty acids fromtriacylglycerols in lipoproteins. Also known as clearing factorlipase, since it removes triacylglycerol from chylomicrons after ameal, resulting in reduction in their size, <strong>and</strong> clearing <strong>of</strong> the milkyappearance <strong>of</strong> chylomicron-rich plasma.lipectomy Surgical removal <strong>of</strong> subcutaneous fat.lipidema Condition in which fat deposits accumulate in the lowerextremities, from hips to ankles, with tenderness <strong>of</strong> the affectedparts.lipids (also sometimes lipides, lipins) A general term for fats <strong>and</strong>oils (chemically triacylglycerols), waxes, phospholipids,steroids <strong>and</strong> terpenes.Their common property is insolubility inwater <strong>and</strong> solubility in hydrocarbons, chlor<strong>of</strong>orm <strong>and</strong> alcohols.Fats are solid at room temperature, while oils are liquids.Non-saponifiable lipids are not hydrolysed by treatment withsodium or potassium hydroxide <strong>and</strong> therefore cannot be

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