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Benders'dictionary of nutrition and food technology

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372saturated fatty acid. The third hydroxyl group is esterified tophosphate <strong>and</strong> one <strong>of</strong> a number <strong>of</strong> water-soluble compounds,including serine (phosphatidylserine), ethanolamine (phosphatidylethanolamine),choline (phosphatidylcholine, alsoknown as lecithin) <strong>and</strong> inositol (phosphatidylinositol).Cell membranes are a double layer <strong>of</strong> phospholipids with thefatty acid side-chains on the inside. The water-soluble compoundesterified to the phosphate interacts with water.This is why phospholipidscan be used to emulsify oils <strong>and</strong> fats in water <strong>and</strong> arecommonly used in <strong>food</strong> manufacture as emulsifiers. From theenergy point <strong>of</strong> view they can be regarded as being equivalentto simple fats (triacylglycerols); they also provide a source <strong>of</strong>choline <strong>and</strong> inositol, neither <strong>of</strong> which is a dietary essential.phosphoproteins Proteins containing phosphate, other than asnucleic acids (nucleoproteins) or phospholipids (lipoproteins),e.g. casein from milk, ovovitellin from egg yolk.phosphoric acid May be one <strong>of</strong> three types, orthophosphoric acid(H 3 PO 4 ), metaphosphoric acid (HPO 3 ) or pyrophosphoric acid(H 4 P 2 O 7 ). Orthophosphoric acid <strong>and</strong> its salts are E-338–341, usedas acidity regulators <strong>and</strong> in acid-fruit-flavoured beverages suchas lemonade.phosphorus An essential element, occurring in tissues <strong>and</strong> <strong>food</strong>sas phosphate (salts <strong>of</strong> phosphoric acid), phospholipids <strong>and</strong>phosphoproteins. In the body most (80%) is present in the skeleton<strong>and</strong> teeth as hydroxyapatite; the remainder is in the phospholipids<strong>of</strong> cell membranes, in nucleic acids <strong>and</strong> in a variety<strong>of</strong> metabolic intermediates, including atp.The parathyroid gl<strong>and</strong>s (see parathyroid hormone) controlthe concentration <strong>of</strong> phosphate in the blood, mainly by modifyingits excretion in the urine. Human dietary needs (about 1.3 gper day) are almost always met; deficiency rarely occurs in adults.The calcium :phosphate ratio <strong>of</strong> infant <strong>food</strong>s is, however,important.Phosphate deficiency is common in livestock <strong>and</strong> gives rise toosteomalacia (also known as sweeny or creeping sickness).Phosphate is also essential for plant growth, hence the use <strong>of</strong>inorganic phosphate or bone meal as fertiliser.phosphorylase Enzyme (EC 2.4.1.1) responsible for the breakdown<strong>of</strong> glycogen <strong>and</strong> starch to glucose 1-phosphate. In vegetablesassociated with formation <strong>of</strong> sugars during ripening <strong>and</strong> onstorage. Important in potatoes during storage because it remainsactive at low temperatures whereas enzymes <strong>of</strong> glycolysis arecold labile, so that sugars accumulate rather than being utilised.photon absorptiometry Technique for determination <strong>of</strong> bonedensity, as an index <strong>of</strong> calcium <strong>and</strong> vitamin d status.

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